I4 SMITHSONIAN MISCELLANEOUS COLLECTIONS VOL. 134 
posterior portion. There is a small posterior cusp placed buccally and 
a brief shelf posterolingually, but without a deuterocone. P® is three 
rooted with a small anterior cuspule and a large tritocone not as high 
as the primary cusp. The deuterocone, though damaged, is seen to 
be well developed and about median in position. In P* the small an- 
terior cusp is higher or nearer the apex of the primary cusp and the 
tritocone is about as large as the primary cusp but more bladelike in 
appearance. The talon carries a well-developed crestlike deuterocone 
and is anteroposteriorly broader and more anterior in position than that 
of P*. M?’, the carnassial, has been converted entirely to a shearing 
blade. This tooth is rather worn lingually, but it is apparent that the 
anterointernal root did not support a talon or deuterocone. M? is a 
small peglike tooth anterolingual to the posterior margin of M?. 
An interesting feature of the Patriofelis upper dentition is the ex- 
tent to which it is carried outward and posteriorly on the strong 
zygoma as in cats and hyenas, and the fossil form, Hyaenodon—quite 
unlike the bears, and possibly dogs. Also, as noted by Matthew (1909), 
the upper cheek teeth are inclined inward, very much so in the 
posterior part of the series, so that the shearing surface, particularly 
of M?, is vertical. The outer wall of M? is nearly horizontal and the 
angle that it makes with the shearing surface is, in consequence, not 
particularly acute. Also, as a result of this, the vestigial M* is im- 
planted at an angle, dorsal to the crown of M7’, so that it is almost 
concealed in the ventral view. Its small crown scarcely reaches the 
plane of shear on M, immediately ventral to it. 
Mandible—The Patriofelis jaws (pl. 4), as Matthew (1909) has 
noted, are deep and massive. The symphysis is deep and elongate, 
and although Matthew considered it as not co-ossified, I find the two 
rami are firmly united. When it was necessary to separate the jaws 
so as to remove them from the skull during preparation, breakage for 
the most part occurred to one side through the canine alveolus. The 
anterior margin of the symphysis is a little less abrupt than Matthew 
has shown for P. ferox, curving gently into the comparatively straight 
lower margin of the horizontal ramus. The depth of the jaw be- 
neath the tooth row is nearly uniform, though a little shallower be- 
neath the posterior root of P, as shown for P. ferox. The mental 
foramina are variable and on the right side two are located one above 
the other beneath the anterior part of P., and two side by side be- 
neath the posterior root of P; and the anterior root of Py. On the 
left side two slightly larger foramina are located, one beneath P, and 
the other beneath P;. There is also an irregular display of foramina 
beneath the incisors close to the anterior margin of the symphysis. 
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