190 MR. JOHN MIERS ON G(ETZIA AND ESPADEA. 
generally remote from the hilum. Even where the fruit is baccate, or where it is more 
rarely drupaceous, as in Brugmansia proper*, or in other cases, as in the Cestrinee, 
Metternicheæ, and Fabianee t, where the seeds are fewer, the nearly straight embryo, 
generally heterotropous, is always imbedded in albumen, with a long terete radicle 
pointing to the basal extremity. Finally, Espadea is irreconcilably discordant with the 
Solanacee, in having the cells of its ovary antieal and postical, not right and left of 
the axis of inflorescence. 
Among other monopetalous families, the Convolvulacec present a few points of resem- 
blance. Wilsonia, the only gamosepalous (and indeed a doubtful) genus of that family, 
agrees with Æspadea in its tubular 5-toothed calyx, its tubular corolla, its bilocular 
ovary, with two erect ovules in each cell; but it differs in its very dissimilar habit, in the 
total absence of a disk, and in its deeply bifid style: the fruit of Wilsonia is unknown. 
Espadea coincides with the tribe Argyrieæ (especially with Maripa) in its bilocular 
ovary, with two collateral ovules in each cell, in its style and stigma, and its drupa- 
ceous fruit; but it differs in its more erect habit, in its solitary extraaxillary ebrac- 
teated flowers, its gamosepalous calyx, its fleshy gibbous corolla, with valvate æstivation, 
in its exalbuminous seeds, with large fleshy cotyledons, and a minute radicle. With the 
other tribes of the Convolvulacee there exists a much smaller degree of analogy. Espa- 
dea differs also from that family, in the same manner as from Solanacee, in the position 
of its carpels. 
With the Zcacinaceæ these genera accord in their ovary partially imbedded in a large 
hypogynous disk, and in their collateral ovules; but these last are erect (not suspended); 
they differ, moreover, in the position of their carpels; in their calyx, corolla, fruit, and 
seeds, which are suberect, without any albumen, having large thick cotyledons and a 
minute inferior radicle. 
If we look to the Logamiacee, we find that Gertnera alone presents a few points of 
similar structure, especially in its bilocular drupaceous fruit, with a single erect seed in 
each cell; it differs, however, from the genera under consideration in its large vaginant 
stipules, its opposite leaves, its panicular or capitate inflorescence, its bracteolate calyx, 
its much shorter stamens, the want of a large hypogynous disk, its seeds having a much 
smaller embryo, with short cotyledons, imbedded in copious cartilaginous albumen. 
Tesserandra in Oleaceæ agrees in having a bilocular ovary, partly imbedded in a 
fleshy torus, each cell having two collateral ovules ; but it differs in its opposite leaves, 
its terminal panicular inflorescence, its corolla, its small extrorse stamens, its style, its 
nucumentaceous fruit, and its superior radicle. 
In Myoporacee we find some genera which agree tolerably. well with Espadea in habit, 
extraaxillary solitary flowers, a similarly shaped corolla, and a bilocular ovary with 
collateral ovules ; but they differ in their deeply partite calyx, their didynamous stamens, 
their peculiar anthers, the absence of a large disk, a different organization of their ovary, 
in their suspended ovules, their monopyrenous fruit, with two or more cells, their sus- 
pended seeds, with an embryo having a long terete radicle enclosed in albumen. 
There is some resemblance in the habit of these plants and in the peculiar venation 
* Ill. S. Am. pl. 56. : + Ibid. pls. 14, 15, 16, 17, 46-51, 59, 60. 
