602 DR. M. T. MASTERS ON THE PASSIFLORAUu rz. 
ovules. In the beginning the ovary is sessile; but in most of the genera of the order it 
becomes, after a time, stalked by the gradual elongation of the central portion of the 
receptacle, which pushes up not only the pistil, but also the stamens, and has hence 
received the name of gynandrophore, or gynophore. In Passiflora multiflora the ovary 
is sessile. The minor peculiarities of the ovary do not call for special comment. 
The processes often met with in the lower part of the gynandrophore demand passing 
notice, inasmuch as they, in conjunction with one of the innermost rings of the corona 
(the operculum), frequently separate, as by a diaphragm, the true nectary, or honey- 
secreting surface of the flower-tube, from the upper portion. 
The species included in the section Granadilla of the genus Passiflora are the most 
remarkable for these processes of the gynandrophore, which there form a kind of pulley- 
like projection, into the groove of which fits the lowest ring of the corona. In other 
species, as in P. elliptica, the same purpose seems fulfilled by a spindle-shaped dilatation 
in, or below, the centre of the stalk of the pistil, and which is covered with fine hairs. 
The styles, usually three in number, project from the apex of the ovary; but in some 
of the Passiflora (sect. Astrophea) the ovary is somewhat cuboidal in outline, flattened 
at the top, and the styles project from the angles at the top of the ovary. 
In Malesherbie the styles project from the dorsal surface of the ovary at some distance 
from the summit, and thus recall the corniculus or horn-like projection from the backs 
of the sepals. 
The genus Barteria is remarkable for the connation of its styles, which thus form an 
undivided column surrounded by a large mushroom-shaped stigma. The stigmatic ends 
of the styles are usually large, more or less reniform or three-lobed. 
The fruit is usually more or less baccate, but is in some genera capsular and dehiscent 
by three valves. No special comment appears necessary, except to point attention to 
the curious manner in which the capsule opens in the Malesherbiæ. In this suborder 
the seed-vessel splits septicidally into three valves, each valve bearing a style or a rem- 
nant of that organ at some distance below the summit of the dorsal surface. Ultimately 
the valves split at the top along the dorsal sutures, the separation extending as far as 
the base of the style. The result is that in the mature fruit there are three valves, 
each valve cleft at the summit, and bearing the persistent style in the central notch. 
At the base of the fruit or of its stipes are the persistent, withered remnants of the 
flower. 
Ovules.—These are generally indefinite in number, horizontal, anatropal, and funicu- 
late, with two coats and a central nucleus. The funicle is transversely articulated above 
the middle; and from the joint proceeds, subsequently to fertilization, an arillus, which 
forms a pulpy investment growing from below upwards till it covers the seed. The 
ovules are generally arranged in four, rarely in two or six, vertical ranks along the 
placentas. | 
Seeds.—The seeds are usually ovoid, obcordate, or heart-shaped, enclosed within the 
pulpy aril just mentioned. The testa is hard and bony, often more or less pitted. In 
the centre is a horny albumen surrounding a straight embryo, with flat leafy cotyledons, 
and whose radicle points towards the hilum. 
