DR. M. T. MASTERS ON THE PASSIFLORACEA. 615 
thread to which the anther is attached has been briefly but correctly described by 
Morren. It consists of a mass of densely packed, very slender, elongated and rather 
thick-walled cells, like those of the filament, but smaller, and through the centre of 
which passes a compact bundle of fine spiral vessels (Plate LXV. fig. 10). "These vessels 
are directly continuous with those of the filament; and as they enter the connective of 
the anther they break up into numerous branches, destined for the supply of the anther- 
lobes. The connective, in the first instance, presents itself at the back of the anther, in 
the shape of two rounded fleshy lobes, with a central depression. These consist of globular, 
closely packed cells; the pollen-cavities are somewhat lateral, and leave between them in 
front a portion of untransformed tissue, with a straight edge. The structure of the anther- 
valves themselves I have not sufficiently examined. The fibrous cells are ovoid, and 
contain a much reticulated spiral fibre. | 
Pollen.—The pollen-cells of Passiflora and Tacsonia are beautiful microscopic objects. 
Their form is globose; and they are covered with a membrane which is raised into pro- 
jecting ridges disposed in a netted manner. The grains open by four valves. For a full 
account of their structure, on which I have no observations of my own to relate, I may 
refer to Schleiden's account*. 
Gynophore.—A transverse section of the gynophore below the lowest cup-like ring of 
the corona shows, going from the centre towards the circumference, an epidermis en- 
closing a parenchyma of spheroidal cells, then a ring of vascular tissue, within which is 
more cellular tissue, in which are plunged five bundles of vascular tissue which, on. 
transverse section, show a semilunar arrangement, the concavity being directed inwards. 
These bundles seem to alternate with the angles of a pentagonal mass of vascular tissue 
and elongated cells; quite in the centre is more cellular tissue. Above the cup-like 
corona the gynophore exhibits the same appearances. A transverse section above the 
stamens reveals an epidermis encircling cellular tissue, in which is a broken ring of 
vascular bundles and spiral vessels, within these more cellular tissue and another inter- 
rupted ring of vascular tissue enclosing the central parenchyma. 
Ovary.—A transverse section of the ovary shows the usual epidermis and parenchyma, 
through which latter run spiral vessels and long, thin-walled tubes. The spiral vessels 
ramify in all directions. Within these there is again parenchyma encircling six vascular 
bundles, three opposite to the placentas, three alternate with them, Each of the pla- 
centary bundles divides into a number of smaller fascicles, destined for the supply of the 
ovules, which, in Passiflora, are usually arranged in four rows on the placentas, so that 
the walls of the ovary contain a network of vascular tissue. In other cases there is only 
a double row of ovules to each placenta; indeed, as the fruit ripens, it often happens 
that two rows of ovules on each placenta become obliterated. The styles are traversed 
by spiral vessels, which do not form a ring. ; ; 
Stigma.—The stigmatic tissue consists of long, loosely packed cells, with conical ends 
overlying a mass of ordinary cellular tissue. 
Ovules. Seeds.—But little need be said in addition to what has been before stated 
respecting the development of the ovule. The arillus is at first a mere sheet of oblong 
* Principles of Scientific Botany (ed. Lankester), tab. 4, pp. 597, 598, and 356. 
