Revision of the Holothurioidea. 339 
made up of 21 branches, while a final single branch is quite separated 
from the principal part. A genital pore terminates each distal 
branch, while the proximal portions of the branches fuse to form 
larger subdivisions, lobes, which may be more or less distinct, or 
else all united as the common base of the papilla. The whole papilla 
may be extended posteriorly to a length of 6 mm or 7 mm. 
A reconstruction from serial sections of the male papilla (Pl. 13, 
Fig. 5), demonstrates that the vas deferens (vd), gives off a tubule 
below the base of each subdivision of the papilla. Soon the tubule 
bifurcates to supply each of the distal branches (a, b, c, d, e, f, 9). 
The gonaduct is ciliated. 
Previous to my paper (1909), a subdivided genital papilla, or 
multiple papillae, have been known only in a few of the Elasipoda, 
while there is no record of differentiation in the form of male and 
female genital papilla. A genital papilla has been reported in the 
male only of Thyone aurantiaca, Cucumaria elongata, and Cucumaria 
laevigata (cf. Lupwıg, 1889—1892). 
Distribution of the pedicels. — Table I gives the 
distribution of the pedicels from the 5 radial canals in 10 of the 
smallest, and 2 of the larger individuals, of the collection. 
The youngest stage I sectioned and reconstructed. In most of 
the other small specimens, the muscle sheet was stripped from the 
body-wall in order to count the pedicels more exactly and to deter- 
mine from which side of a given radial canal the pedicel arises. 
All of the specimens have the 10 tentacles, enteric canal, stone- 
canal, Polian vesicle and gonad of the adult. Specimen 1 has in 
all 40 pedicels some being only rudimentary buds and all are radial. 
From the mid-ventral radial canal 4 arise to the right and 7 to the 
left. From the right and left ventral radial canals, 9 and 7, 
respectively, are given off ventrally and none dorsally (Pl. 13, 
Fig. 6). 
From each of the right and left dorsal radial canals, 5 pedicels 
arise ventrally and 1 is terminal posteriorly, while there are none 
dorsally (Pl. 13, Fig. 7). In connection with the older pedicels 
.of this stage ampullae have developed. 
In following Table I, it is to be noted that in specimen 2, with 
51 pedicels, the appendages have begun to appear dorsally from the 
lateral ventral radial canals, but not as yet similarly from the dorsal 
canals. Specimen 3, with 78 pedicels, has 4 pedicels dorsally from 
each lateral dorsal radial canal. Specimens 4 and 5, with 142 and 
Zool. Jahrb. XXIX. Abt. f. Syst. 23 
