146 MR. 5. J. A. SALTER ON A SEXUAL MONSTROSITY 
extended into foliaceous prolongations. Rarely the imperfection in the ventral sutures 
has occurred in the centre of the ovary, and not at its extremity: in such cases the 
ends of the ovary have been closed, and one, two, or three small openings have occurred 
in the middle, according as two or more of the carpels were separated on one side or on 
both (Pl XXIV. fig. 3). In these instances the everted edges bore cellular processes 
containing pollen. All the polleniferous organs connected with these monstrous ovaries 
were either attached to the edges of the carpellary leaves where they were divided, or 
sprung from the placentas. The position which they occupied was that of ovules, and 
both in structure and form they passed by degrees from organs identical with anthers to - 
others which were identical with ovules. I incline very strongly, therefore, to the 
opinion that in all instances they were modified ovules, and that in their earliest con- 
dition—in their incipient development—they commenced as such. 
The forms of these pollen-bearing bodies have been very diverse, but for convenience 
of description, rather than for any other reason, I may mention four different varieties 
in which I have found them. 
First, that of a bilobed anther. This in many instances has been a tolerably perfect imita- 
tion of the genuine anther of the same flower. Such specimens are figured in Pl. XXIV. 
figs. 1 & 2. In all cases these anthers were sessile upon the free edges of the carpellary 
leaves, and placed towards the distal end of the ovary. In structure they were essentially 
the same as true anthers. The external coat consisted of epidermis, or exothecium: the 
internal coat was composed of spiral cells and free spiral fibres, constituting a true endo- 
thecuum. Within the latter was a mass of pollen. Sometimes these antheroid organs 
have been more or less expanded and foliaceous, but essentially of the same structure. 
There is nothing in the form or ultimate anatomy of these bodies to indicate an ovular 
_ nature, and the opinion that they are modified ovules is merely inferential. 
The second form consists of globular or oval bodies attached within the ovarian cavity 
to the placenta by a pedicle more or less constricted. The general contour of these 
bodies suggests a modified ovule, and their position is in keeping with that idea. Their 
structure, however, I have found entirely antheroid, and destitute of the particular 
arrangement of parts constituting an ovule. They have consisted of a chamber filled 
with pollen surrounded by inenchyma (endothecium), and clothed by epidermis (exothe- 
cium). The stalk has been usually destitute of vascular tissue, and composed of paren- 
chyma with some spiral cells. These bodies vary a good deal in form and size, being 
occasionally very large and oval. One of these polleniferous processes I have carefully 
mounted as a microscopical object, and the accompanying figure (Pl. XXIV. fig. 4) 
is a faithful illustration of the specimen magnified about 35 diameters. I could not 
make a section of the little polleniferous body from its minuteness; it was therefore 
necessary to compress it so as to render it sufficiently thin for viewing with transmitted 
light, otherwise the nature of the opake pollen-mass in its centre would not have been 
clearly apparent. This compression has modified the form by pressing out the soft stalk 
and rendering it much broader in proportion to the rest of the organ than it was originally. 
A third form was usually found fairly within the cavity of the ovary, and below the 
free split edges of the carpels. This organ undoubtedly was a modified ovule. It was 
