DR. MOXON ON THE ANATOMY OF ROTATORIA; 459 
the lips of the aperture may be commonly seen partially closing and opening. Starting, 
as stated, from this vesicle, the tubes (fig. 1 û), as usual, seek the mid-lateral region and 
approach the surface, where they aequire some fat-granules in their parietes; here the 
first cilium-funnel is situate (fig. 1): if the side of the Floscularia be towards the 
observer this will not be visible, as I shall presently explain. The tube continues to 
ascend in the side, as high as the neck, where there is another funnel (fig. 1 4), situate 
just behind and below the lateral feeler; it then passes, still ascending, to the back of 
the neck, and crossing the middle line anastomoses with its fellow just behind the faucial 
sphincter ; above the tube, near the middle line, and below and outside the dorsal median 
feeler, is a third funnel, lying horizontally (fig. 1#"); and above and outside this is a 
fourth (fig. 1 X"), hanging vertically. I have been unable to find more; but as the smallest 
number yet known in any Rotifer is five pairs, I have little doubt that the missing one, 
which should be in the side of the neck, will some day be observed. I have plainly seen 
the cilium-funnels in activity in the egg of Floscularia; they may also be seen in the eggs 
of Philodina (fig. 4° k). 
Of the peculiar cireulatory apparatus described by Mr. Gosse* as compensating for the 
supposed absence of the water-vessels in Floscularia, I could never see any trace; but, in 
the contractions of the body of the animal, the granules alluded to by him move freely 
about in the perivisceral space. 
Leydig, in his account of Pterodinat, states that he was unable to see any ciliated 
appendages to the water-vascular system of that Rotifer. "These appendages I found to 
be five in number on each side; their positions are shown in the accompanying drawing 
of Pterodina (fig. 5 k k). | | i 
Now with regard to what I have here called cilium-funnels, a view at present prevails, 
according to which these are described as blind sacs enclosing a “long flickering cilium." 
The grounds on which I venture to differ from this view are, I think, conclusive. The 
Rotifer which is best suited for the observation is Euchlanis dilatata. It is common, and 
of good size. If a large specimen be chosen, and turned on its back (fig. 7°), four pes of 
funnels ean be made out: but two pairs (fig. 7* # k") in the lateral ventral region are 
plainly visible to a low power; these are narrow in outline, and display, 52 beautiful 
degree, the candle-flame-like appearance that is described as a flickering cilium. If the 
animal be now turned on its side (fig. 7), a feat which can be easily managed in aS 
compressorium, these two funnels will be so faintly marked as to be scarcely per ceivable; 
Whilst a. third (fig. 7 k"), situate in the neck, which in the ventral view of the animal 
was barely visible, now shows itself as a flickering cilium. If now the observer search 
carefully, he will be able to discern each of those lateral-ventral tags which in the PN 
View had the candle-flame-like appearance (fig. 7 4 k"), but now m quite — is = 
—that of a broad triangle, with its lower margin well defined, € — dini 
with short cilia in full activity—this not doubtfully or ss the presence of the 
and const i bserver's mind not only oi th 
nstantly, so as fully to satisfy the obse cilium. The tube (fig. 7 i) can be seen 
cilia, but of the total absence of any large | 
descending and opening into the triangle. Whether this is a triangular Nere ampulla, 
* Popular Science Review, No. 2. + Siebold and Kölliker’s Zeitschrift. 
