of Peripatus nove-zealandiz.. 205 
Segmental Organs ?—These are what I previously called 
salivary vessels.” They form a series, on either side, un- 
connected with each other, but running into each leg, with 
the exception of the first three pairs. ach consists of a 
trunk coming out of the leg, which divides into two ; and these 
branches, after several foldings, unite together, thus forming a 
closed loop. They are filled with colourless granulated cells. 
They were regarded by H. Milne-Edwards as nerves passing 
into the legs (Ann. des Sci. Nat. 2° sér. xviii. p. 128*) ; but I 
have carefully dissected some out, and find that they have no 
connexion with the nerve-cord; and I also feel confident that 
they do not open into the body-cavity, Consequently I do not 
feel sure that they should be considered segmental organs. 
Stime-ducts.—These pass from the oral papille along the 
back to about the centre of the body ; they then turn forward 
and throw off branches ; they then turn backward again, and 
reach nearly to the posterior end of the body. They are pro- 
bably homologous with the supposed segmental organs. 
Respiratory System.—I think Mr. Moseley is right in con- 
sidering my “spiral fibres” trachee; but then all resem- 
blance to the trachez of insects vanishes. Professor Ray 
Lankester is probably right in considering that the tracheal 
systems in Pertpatus and in insects have been independently 
developed (Quart. Journ. Microse. Sci., Oct. 1877, p. 439). 
‘Circulatory System.—I have succeeded in dissecting out 
the dorsal vessel; it contains a greenish-yellow fluid. Mr. 
Moseley is certainly incorrect in denying the existence of the 
lateral canals’ of Grube. It is satisfactory to me to think 
that I had demonstrated them before I knew that they had 
been previously described. Whether they belong or not to 
the circulatory system may perhaps be considered an open 
question ; but they contain, especially at the anterior end, a 
yellowish-green fluid like that in the dorsal vessel. 
Reproductive System.—By selecting small individuals I 
have succeeded in finding two males. Mr. Moseley’s descrip- 
tion of the male organs is very accurate; but they lié above 
the alimentary canal, and not below it. With the exception 
of these two specimens, all the rest were what I consider to be 
hermaphrodite. They all had the organs described by me as 
testes ; but in one individual the testis was absent on one ovi- 
duct, but present on the other. In the early spring (September) 
these contained no spermatozoa; but in November they were 
abundant. During all this time the oviducts were crowded with 
* It is astonishing what a very full and accurate knowledge of the 
anatomy of this animal M. Milne-Edwards obtained by the dissection of 
one badly preserved specimen. 
