KNOWLEDGE OE THE MUTATING OENOTHEEAS. 25 



Another race of O. I/amarcMancc, yvhose characters differ only quantitatively from those 

 of DeVries's race, I have studied carefully and compared with the latter. This race was 

 derived from the coast near Liverpool and has already been referred to in connection with 

 the history of 0. ZamarcJdana (see p. 18). Bailey (1907) showed that this form was 

 O. Lamar ckiana, and MacDougal (1907, p. 8) found from cultures that it produced several 

 mutants. I have since made more detailed comparisons of forms from this locality, and 

 have discovered a number of new forms, some of which have been isolated as races, and a 

 few of which will be referred to in this paper. The O. Lamar chiana race from St. Anne's 

 is almost identical with that of DeYries, being much closer tlian the last race described, 

 j<dt there are a few c[uantitative differences in such features as width of rosette-leaves, 

 diameter of rosette, and number of long hairs on the sepals, which appear to be 



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constant. 



I have seen one other race of 0. Lamarchiana from this localitv which differed 

 considerably in the shape of its stem-leaves, which were longer and tapering to a petiole 

 at the base. This race stood somewhat nearer to O. yrandiflora in its characters. One 

 may therefore expect to find races of O. Lamarchiana bridging more or less completely 

 the gap between this species and O. grandiflora. A series of hybrids between certain 

 races of O. Lamarchiana and 0. grandifiora, w bich I am now cultivating in Fg, may throw 

 some light on the origin of such intermediate races. For the Fi contained two types 

 each of which approaches one parent although in certain features resembling the other. 

 The two hybrid races were therefore different from either parent, though intermediate. 



Another very interesting race, which I received from the Bremen Botanical Garden 

 under the name O. Lamarchiana^ Ser., var. cruciata, must be briefly described here. In this 

 culture 53 plants developed. Their rosette-leaves closely resembled 0. Lamarchiana, being 

 deeply crinkled, but frequently broader in the middle of the blade than typical for that 

 form. Many leaves were also of rather larger size than 0. Lamarchiana usually attains, 

 the maximum size being 9'5 X 36 cm. Of these rosettes, 11 showed a striking peculiarity 

 which I have not seen in any other form* (see PL 1. fig. 12), and several others showed 

 traces of it. The leaves were peculiarly curled and sickle-shaped, the blade being almost 

 wholly undeveloped on one side of the midrib. There was no lack of chlorophyll and no 

 evidence of any parasite, although it is possible the condition may be pathological. The 

 peculiarity appears when the seedlings are less than two months old. I have not yet 

 made the test, but it is probable that the peculiarity is at least partially inherited. The 

 only case I have seen at all resembling this is a single rosette (see Gates, 1910, pi. 31) 

 in a culture of an 0. Lamarchiana from Lancashire. In this rosette many of the leaves 



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were green on one side of the midrib and colourless (lacking chloroplasts) on the other. 

 The colourless portion continually died away from lack of nourishment, but there was no 

 curving around of the green half of the leaf. I interpreted that case as a probable 

 sectorial chimera, but it is not probable that that explanation could apply in the preseut 

 case. 



The flowers of this race also presented several remarkable peculiarities. The style was 



* I have since observed similar aberrations ia ceitaia other races. 



SECOND SERIES. — BOTANY, VOL. VIII. 



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