239 



VIII. A ContribuMott to the Skicly of the Evohition of the Flower, with 



spec 



reference to the Hamamelidaceae, Caprifoliaceae, mid Cornacese. By Aetiiur 

 Samuel Horne, ^.^6'., F.L.S., F.G.S., Lecturer on Botuny in the Vwyal 

 Morticultural Society's School of Horticulture. 



(Plates 28-30 and 13 Text-figures.) 



Eead 4th December, 1913. 



^ H 



Contents. 



1. Introduction , , 



2. Hamamelidaceai 



3. Caprifoliacege • . 



4. Cornacese .... 



(a) Cornns 



(b) Alangmm and Marlea 



(c) Aucnha 



1 



(d) Garrya , 



(e) Helwingia 



(f) CoroJcia 



(g) Melanophylla 



(h) GriseHnia 



(i) Nyssa 



5, Final Discussion , 



Argument : — The Interpretation of ^Resemblances and Differences. 

 Origins. Phylogeny of the Hamamelidacese. Phylogeny of the 

 . Caprifoliaceae. Phylogeny of the Cornacese. The Vascular 

 Structure of the Flower in Angiosperms. The Ovule in Angio- 

 sperms. Ovular connections. Ovular Vascular System. The 

 Phylogeny of Angiosperms 291-302 



Page 

 239 

 245 

 2.51 



2G1 

 261 



265 

 269 

 274 



278 

 282 

 284 

 284 



287 

 291 



1. INTE.ODUCTION. 



xHE structure of the flower of Bavidia, a genus referred to the Cornaceae in 'Die 

 Natiirliche Pflanzenfamiiien,' has been ah'eady described by the author* in a paper 

 deahng with the structure and affinities of Bavidia involucrata, published in the 

 'Transactions of the Linnean Society,' in 1909. During the study of Bocidia there 

 was occasion to examine the flowers of several o^enera of Cornaceae, and in so doing 



o 



striking structural differences were discovered not only between representative genera 

 of the subfamilies but also between the genera tliat have been grouped in a particular 

 subfamily, such as, for instance, the Cornoideae. At the same time, as reference had 



* A. S. Home in Linn. Soc. Trans. 2nd ser., Bot. vii. (1909) 303-326, t. 31-33. 



SECO>^D SERIES. — BOTANY, VOL. VIII. 



2o 



