250 



ME. A. S. HOENE — A CO^'TEIBUTION TO 



duced stamens. OHt 



nectaries as r* 



Hustigma repla 



was ohserYed by the writer i 



in 1909. The normal flowe 



petal 



states that the "didynamous squamae 



j> 



]n 



An instance of the transfor 



of petals into stamen 



flowers of Corylopsis sjncata gathered from Kew plants 



P 



fiT 



petals and five stamens 



A 



transitional series occurred between the normal and the entirely apetalous fl 



plete 

 The 



transitional flowers possessed foliose stamens, each consisting of a 



broad filame 



d rudimentary 



lobes 



. The genus Fotliergilla possesses a number of stamens 

 These emerge from a perigynous collar, which surrounds 



arranged in a single verticil, 

 the ovary. The writer finds fi 



Baillon f investigated the development of the stamens and found that they rep 



seventeen 



enty-two in F. Gajxleni and maj 



al distin 



S 



Fofhe 



eally transformed sepals and petal 



apetalous, it is possible that the outer 



The course of the vascular tissue has been followed in four g 



JJamamel 



Fothergilla, Corylopsis, and Fm 



The vascular cylinder (Text-fig. 1, A 1) separates 



the base of the ovary, into an outer and an inner series of bundles CD 



of the c 



ridge of 



series branch just bel 

 e, to form externally 



the level of the insertion of 



lie stemonal and perianth 

 pellary bundles, d and I ( 



B 



The bundles 

 3 perigynous 

 and pe; and 



The former 



internally the dorsal and lateral ca 



then leave the ovary. The inner bundles vary in number and disposition in differ 



genera 



I 



Foth 



C 



jrgilla, two bundles are present in the septum as shown in 

 {Farrotia), and each divides before leaving it to form two, which traverse the corres- 

 ponding placentae, B 3. Thus the bundle m (C 1) forms m 1 and m 2, which traverse pi 1 



and pi 



pectively (B3J. 



A single branch from m 1 supplies an ovule in the left 

 hand loculus and one from m 2 an ovule in the right-hand loculus. But in Hamamelh 

 Zuccariniana, where the loculi are uniovulate, only one placenta corresponding to ^H 



and pi 2 is found, and the 



ptal bundles, m, p 



ched (A 3, A 4) 



In 



this case, one branch passes to the ovule from the bundle, m. 



The above-described system is of a similar type to that which is found in the gem 

 Saxifraga. It resembles in some details that of Cmyiniis hetulus, described by Benson 

 and Welsford. 



Summary. 



1. A complete series from hypo 



to 



epi 



forms of flower exists in the 



Haraamelidacea?, ranging from Fisiylium, by way of the perigynous genera JIamameUs, 

 Corylopsis, and TricJwcladus to Eustigr,ia. Another series ranging from a FislyUum-Yike 



Of 



n 



y^j 



form is only represented by Sycopsis, a genus which simulates epi 



fact that the ovary is invested by a closely adpressed perigynous tube. Loropetalum 



epigynous, but the origin of this condition needs investigation. 



Bhodolela, according to published descriptions, exhibits variability 



2. The genus 



the mternal structure of the ovary. 



D. Oliver in Trans. Linn. Soc, Bofc. xxili. (1860) 83. 

 t Benson and Welsford in Ann. of Bot, xsiii. (1909) 630. 



t H. Baillon in Adansonia, x. (1S71) 1^7 



