THE 



rLOWER. 251 



3. The ovules in Ramamelis, Fothergilla, Corylopsis, and Farrotia are suspended from 

 parietal placental that are situated immediately above the septum in each. 



4. The Hamamelidaceye exhibit a complete series in the reduction of the number of 

 ovules, ranging from Liquidamhar, with numerous ovules in each loculus, by way of 

 Disanthus, in which Baillon records three ovules in each series, and Father gilla. 



CorylopsiSy and Pa^ 



two ovules (one abortive) in each loculus, as in several 



Araliaceee and Umbelliferae, to F[ama7nelis, in which each loculus is uniovulate. 



5. According to Baillon, the ovaries of Ilamanielis and Corylopsis are often arrested 

 in development and remain small and sterile. This feature is of special interest, owing 

 to the fact that genera with polygamous and with unisexual flowers are frequent in 



the Hamanielidaceae. 



6. The ovules of the Hamamelidacese possess a bulky nucellus which becomes invested 

 by two integuments and resembles in structure that of the Saxifragaceae and Rosaceaj. 



7. The bracts of Farrotia are tripartite and correspond morphologically to a stipulate 

 leaf. One or other of the segments becomes specially developed and conspicuous. 

 Hence the conspicuous portion may be homologous with either a stipule or the lamina 



of a leaf. 



8. A complete series in the metamorphosis of petals into stamens has been observed 



in Corylopsis spicata. 



9. The course of the vascular tissue in the flower has been followed in four genera 

 Hamamelis, Fothergilla, Corylopsis, and Farrotia. The same type prevails in them all 

 and is similar to that which is found in Saxifraga. 



3. CAPRirOLIACEiE. 



The general details of floral organization in the Caprifoliacese are so well known that 

 they need not be recapitulated. Hence the morphology and vascular structure of the 

 ovary and ovule will receive special consideration. 



Morpliology of the Ocary. 

 The ovary of Leycesteria, the type genus of the family, is five-chambered below, 

 owing to the fact that the margins of the carpels are in this region congenitally united, 

 forming a solid axial portion with radiating septa (Text-fig. 2, Al), but the margins 

 are free above, so that the accretion is incomplete (A 2). Hence the biseriate ovules are 

 axially or parietally borne according to their situation. The fruit contains many seeds. 

 Lonicera possesses a trilocular ovary, wliich is structurally similar to that oi Leycesteria 

 (B 1-B 3), but although the ovary is pluriovulate, few seeds are formed. Frequently in 

 X. Caprifolium only one seed reaches maturity, the septa being ruptured or pushed 

 aside by the developing seed, which eventually occupies the whole cavity of the ovary. 

 Here the ovary does not undergo structural modification during growth in connection 

 with the habit of producing few^ seeds. Abelia and Linnma also posses.s trilocular ovaries 

 of the Zeycesferia type, but in each case one loculus is uniovulate, the ovule being 



