278 MR. A. S. HOENE — A CONTEIBUTION TO 



Myrica and the exalbuminous seeds of Salix, Jnglans, JBetida, and Myrica. The ovules 

 of Garrya are anatropous, and the seeds contain copious endosperm. 



Again, since the embryology of Garrya is not yet understood, it would be wise 

 to reserve a final judgment until a more thorough comparison with the Araentiferse 

 could be made, the embryology in the latter group being already known from the work 



of Benson and others. 



Now genera with amentaceous inflorescences and dioecious, apetalous, hypogynous 



flowers occur also among albuminous families with anatropous ovules, and their ovaries 



are comparable in many cases to those of Garrya. Thus the resemblance to the Ilama- 



melidacese is comparatively close. The structure of the ovule forms the only considerable 



difference. Garrya, in many features, is not unlike Soepay now incorporated with 



Aporosa, a genus of the Phyllantheae ; but the Phyllanthese are defined as possessing 



ovules with a ventral raphe, which is not the case in Garrya. Lastly, Garrya may be 



favourably compared with Forstiera, a genus of Oleacese. 



{e) Helwixgia. 



Morphology of the Mower and Ovule. 



The flowers are unisexual and seated upon the midrib of a foliose bract. The male 

 inflorescence consists of a group of separate pedicellate flowers, found to be either 

 trimerous or tetramerous, both kinds being present in the same inflorescence. Each 

 flower possesses a polyphyllous perianth and a single whorl of stamens, which alternate 

 with the perianth leaves. The female inflorescence comprises a single, minutely pedicel- 

 late, epigynous flower, which bears at its base a small foliole. The flower consists of an 

 inferior polycarpellary ovary terminated by free styles and surmounted by a single whorl 

 of perianth leaves, resembling those of the male flowers, which alternate with the stigmas. 

 Stamens, or the rudiments thereof, are entirely absent. Both whorls are usually tetra- 

 or pentamerous, but in one exceptional case a pentamerous ovary was surmounted by a 

 tetramerous perianth. The ovary is multilocular, each loculus containing a single, 

 terminal, axially suspended, descending, anatropous ovule with abaxial raphe. The 

 ovule is radially orientated, and since the nucellus of the mature ovule rests between 

 the axial portion of the ovary and the abaxial raphe, the orientation is the inverse of 

 that characteristic of the Araliacese, where the ovules are also radially orientated. 



Owning to its radial orientation, vertical sections of the ovule are easily obtained. 

 Such a section of the youngest ovule found is shown in PL 29. fig. 73. The 

 integument shows no sign of dual origin. The nucellus approximates closely to 

 the condition in lAlium, and consists of three short axial rows made up each 

 of two or tliree cells. PI. 29. fig. 73 shows a single enlarged subepidermal archesporial 

 cell, of which the subsequent development has not been traced. The jiucellus soon 

 assumes the vertical position, owing to the activity of meristem at its base, and by the 

 time the embryo-sac is formed the ovule has elongated considerably (Text-fig. 13, B). 



