290 MR. A. S. HOENE— A CONTEIBUTION TO 



the characters of both parents, viz., G. elliptica in which the flowers occur in 

 axillary trimerous groups, and G. Fadyenii in which they are situated singly 

 in the axil of each bract. 



d. Rudiments of a single whorl of stamens in the female flowers of Griselinia 



litt oralis and lucida and of the absent petals in G. lucida. 



e. Rudimentary terminal parietal placentge in Corokia Cotoneaster. 



2. The terminal ovules are suspended axially in JDavidia, Cornus, Helwingia, and in 

 bilocular forms of Marlea and Corokia; probably axially by origin in Nyssa and 

 Griselinia, and parietally borne in Aucuba and Garry a. 



3. The ovule exhibits considerable variety in organization : 



a. It is in IVyssa and Davidia of the dorsal type (raphe ventral or adaxial) with 



radial orientation ; of the ventral type in Relwingia and Corokia (raphe dorsal 



the inverse of the former : and of the 



abaxial) with radial orientation, but 



ventral type with tangential orientation in bilocular forms of Cornus and 

 Marlea. 



b. The nucellus in the genera investigated (including Davidia and JSTpsa described 



as bitegumentary by Harms and Wangerin) becomes invested with 



3 



gument 



c. The integument is completely free from the nucellus in Aucuba and Griselinia, 



but in the specialized ovules of Davidia and Belwingia is only found as a 

 separate organ towards the summit of the nucellus. 



d. The nucellus consists of about seven axial rows of cells in Aucuba; four or five 



m Cornus^ Davidia, Griselinia, and Garrya ; and three in Belwingia, where it 

 approximates to the condition obtaining in LiUum and Inula. 



e. The nucellus of Aucuba, in the formation of a definite layer of parietal tissue, a 



small epidermal cap, and in subsequent bulk-extension owing to cell-division, 

 approaches the type described by Pechoutre for the Eosaceie. 

 / The germination of more than one megaspore, already recorded by the writer 



for Davidia, occurs also, but not to the same extent, in Aucuba. 



4. The floral vascular systems of the Cornaceae do not conform to one type as in the 

 Caprifoliacese. It is in Cornus of the Caprifoliacean type. The systems of Griselinia, 

 JVyssa,and Helwingia exhibit Araliacean tendencies; those of Davidia and Marlea are 

 transitional in character between that found in Aralia and the synthetic systems of 

 Cornus and the Caprifoliaceaft. The stage of economical advance reached by Viburnum 

 has not been attained. 



a. Infra-locular extensions of vascular tissue occur in Aucuba and unilocular forms 



of Marlea. 



b. The ovular trace is anomalous in Griselinia, Nyssa, and unilocular forms of 



Corokia. 



c. Branches from the ovular trace traverse the peripheral region of the integument 



in Davidia and Corokia. 



