208 MONOGRAPH OY THE LABOULBENIACE^. 
nil cases tlie primary appendage is originally a terminal structure, developed at least 
in part, often entirely, from the terminal spore segment, which is as a rule distinctly 
smaller than the basal, and never under any circumstances concerned in producing any 
portion of the true receptacle. While this terminal character of the primary appcn- 
da«'e is often obscured, asinZodiomyces, Rhachomyces and other genera, it is in many 
cases sufficiently evident ; for example in such instances as Stigmatomyces (Plate 
figs. 2-13), Cantharomyces (Plate VII), and the like. In the genus Ehachoniyces 
the primary appendage is represented by the single small bristle at the very base of 
the whole series of appendages, which are thus nearly all secondary and quite different 
in origin from the originally terminal primary one. Again, in Zodiomyces the origi- 
nally terminal primary appendage is soon sloughed off, its place being taken by a 
multitude of secondary ones, also quite different in origin. 
Although they are very important frotn a systematic point of view. It would be 
r 
quite superfluous in the present connection to enter into any detailed description of 
the many variations of form and structure which the appendages, both primary and 
secondary, exhibit ; and further details should be sought in the special descriptions of 
the genera. It is sufficient for the present purpose to say that the appendages, using 
m 
include both primary and secondary, may be solitary or very 
tly elongated or consisting merely of single short cells; simple or branched 
t variety of ways, either sympodially or monopodiallv, or more or less irreo-ul 
■h 
hyaline or deeply colored ; stiff and bristle-like or flexuous and slender ; In short, as 
may be seen by a glance at the accompanying plates, showing wide variations as 
to 
details even in nearly related 
The sterile appendages, or the sterile portions if such are present of fertile appen- 
dages, which are always more or less filamentous and composed of successive cells 
placed end to end, may become very highly developed, forming a dense tuft which is 
often much more conspicuous than the main body of the plant itself. The function of 
hly developed sterile branches is doubtless primarily that of protection for the 
^y> V 
delicate trichogyne, which is subsequently developed, and the> 
further office in facilitating the fertilization of this organ by holding around it a drop 
of water, which is usually found at times condensed on the surface of the hosts, the 
ty of which are apt to hide by day 
It mig 
be supposed that since tliese sterile portions of the appendages extend, in many cf 
beyond or around the tip of the perithecium, that they were of some assistance 
the dissemination of the spores; but it is a curious fact that the latter, despite th 
generally viscous nature, are only in exceptional cases found adherent to any port! 
/ 
