218 
MOXOGRxiPH OF THE LABOULBENIACE^. 
pi 
ariably 
division of the basal cell, never in any case from tlie terminal cell, whe 
cnt. Although the products of the division of the terminal cell are in^ 
or male, it is not true, as might be supposed, that the basal cell or its derivatives ha^ 
any inlierent female character; since, in many cases, both normal and abnormal ai 
theridia and antheridial branches may arise below the point of insertion of the fema 
organ, or even, in exceptional cases, replace it entirely (Plate II, figs. 7-8). Whil 
then, the primary appendage is, as a rule, terminal, the 
■y 
p 
folio 
^y 
as Avell as the 
ption of Amor- 
This 
phomyces, where both are developed terminally from the unsegmented spore, 
lateral origin is, however^ very often obscured in the fully developed plant, from the 
fact that the perltheciumj as it develops, is apt to push the appendage more or less to 
one side, and assume an apparently terminal position ; as, for example, in the genus 
F 
K 
Stigmatomyces or in Laboulbenia. 
We 
ady seen that the basal 

f the 
germm 
s 
pore begins 
development by the formation of a cross partition which divides it into an upper and 
a lower cell, and that the base of the latter becomes modified to form the foot. 
The 
development of the upper, although varying considerably in the different genera, may, 
perhaps, be best illustrated by reference to the series of figures (Plate I, figs. 1-24) of 
Stigmatomv 
IS 
pical of the more 
form 
In this series, fig. 2 
epresents the spore after it has become' attached 
to the host, the foot Is beg 
form and the upper half is considerably enlarged. Figs. 3-5 illustrate the furth 
development of the upper cell, the lower still remainino- as at fi 
I 
b 
become divided into two superposed 
fig. 6 
f which (b) forms 
of 
append 
fo 
ige. The lower c 
r 
basal cell and the 
divisions is not apparently' 
happens that the partition 
that which separates (a) fro 
(?/) then divides in two by 
b-basal cell (a). This scqi 
f 
o 
has formed 
It, and in other forms, at least, it more frequently 
separates (a) from the basal cell is formed before 
In fig. 7, the nucleus of cell (a) has already divided, 
bee 
daug 
nuclei. In fisr. 8 
formed and the cell (a) has become divided into an 
O 
d 
of the append 
change as the 
develop into t 
(h) remaining permanently 
pper 
fur 
lis wall has 
d a lower 
IS the base 
Of th 
(a' and a"), the lower remains without fui 
fem 
of the receptacle 
the upper («') 
■gan and the perithecium. It will be noticed that 
early a stage as is represented in fig. 8, the antheridial cells have beg 
mature and to discharge their antherozoids 
Cell (a) next begins to grow upward and 
I 
^ 
