220 MONOGRArH OF TEE LABOULBEXIACE^. 
different specimens, but is usually not greater than is represented in fig. 15 {it 
and in fig. 16. Before fertilization it can be clearly seen, as in the last-mentioned 
figure, that tlie trichogyne and the basal cell {c) constitute a single cell, and the 
nucleus is usually readily made out in the position indicated at the base of the 
tie. Meantime the cells [n n) and {o o) of fig. 13 have increased in size, and 
former have betrun to ^row up still further around the base of cell ie), fig. 13, (t"), 
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fig. 14 
With the maturity of the trichogyne the female organ has completed its develop- 
ment, and we have the condition represented in fig. 15, from which it will be seen that 
the latter, which may be conveniently termed the procarjn, consists of three distinct 
parts, which, so far as is known, are present in all genera of Laboulbeniaceo8. Of 
these the uppermost may be properly called the tricliogyne, and in the present illustra- 
tion is unicellular, though often, as will be seen presently, far more complicated in 
structure ; the middle 2)ortion in this, as in all other instances, also unicellular, which 
we may call the irichophoric cell (ie"), and the low^est portion (/), which may be termed 
the carpogcuic cell, being that portion of the procarpe which is fertilized, and the only 
part which persists and undergoes further development. As is shown in the figure, 
the carpogenic cell is completely surrounded by eight cells, four of which (o o) lie be- 
low and around it, and, remaining undivided, form the basal cells of the mature 
perlthecium ; while the four others [n n) completely surrounded it, and, as will be seen 
later, form by further division the wall- and lip-cells of the perithecium. These 
eight cells are arranged in four longitudinal rows, and, in the stage represented, the 
upper four have already begun to grow up around the trichophoric cell, the lower 
half of which is now enclosed by their advancing tips. In the stage represented in 
fig. 15, the antherozoids, which have been continuously escaping from the antheridia 
since the stage represented in fig. v was reached, begin to adhere to the trichogyne, 
T 
often in larger numbers than are represented in the figure, and fertilization is accom- 
plished, probably with considerable rapidity if one may judge from the rarity of 
conditions which are intermediate between that represented in fig. 15 and that shown 
in fig. 19, where the trichogyne has entirely disappeared. In the comparatively in- 
frequent instances in which one finds an adherent trichoixvnc belomriucr to a T)i'ocarpe 
ni which fertihzation has evidently been accomplished, slight elevations may be seen 
which coincide with the position of usually more than one of the antherozoids (figs. 17 
and 18). In such cases, althougli it is very difficult to determine the actual presence 
of conjugation in the case used for illustration, it is evident that a wall has been 
formed around the autherozoid, which often seems inflated and nearly empty of 
contents. 
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