222 MONOGRAPH OF THE LABOULBENIACE^. 
sterile, while the others constitute the ascogenic cells, and at once begin to bud up- 
ivard:'tlie buds (ficr. 21, as) developing into asci (fig. 22, as), and in this, as in a 
majority of cases, arising in a more or less distinctly double row. (See Plate I, fig 
38 ; Plate III, fig. 1 ; Plate V, fig. 18.) 
In Stigniatoinyces the four ascogenic cells, but two of which appear In the figures 
are at first symmetrical neither in form, size, nor arrangement; but, as the asc 
boijcin to develop, become so placed 
posterior, and 
on either side. In fig. 23, which represents an antero-posterior view, the two 
lateral ascogenic cells are shown, placed more or less symmetrically w^ith reference 
to one another, the anterior and poste 
cells (not shown in the optical 
section) occupying a similar relative position in front of and behind them. • 
Eeturning now to the perithecium proper, which we left in the condition repre- 
sented in fig. 15, it will be remembered that it originftted as a single cell (fig. 10, c), 
which has divided several times, and that the upper products of these divisions have 
p^rown up around the base of cell {d), from which, as we have seen, the female organ is 
developed. In this stage (fig. 15), it will be seen to consist of the stalk-cell {p), the 
secondary stalk-cell (Ji), and three basal cells (o), but two of which are visible in the 
figure, and four primary wall-cells {n,n), which surround the carpogonium (/) and the 
base of the tricliophoric cell (e"). At a stage slightly earlier than that represented in 
fig. 15 a fiu-thcr development from the three basal cells takes place, which is not indi- 
r 
cated in the optical section. This development consists in the upgrowth from the basal 
cells (o) of four cells corresponding to the wall-cells, but alternating^ with them and 
o 
lying partly between them and the carpogonium. As they continue to grow upward 
and to increase in size, they separate the wall-cells completely from the structures de- 
veloped from the carpogenic cell, growing up around the latter in a fashion exactly 
resembling that of the wall-cells. There are thus developed from the three basal 
cells, eight cells arranged in two series; an outer, the primary wall-cells, four in num- 
ber; and an inner, also consisting of four cells. The further growth and the succes- 
sive divisions of the cells of these two series, although its course is identical, is, 
quite mdep 
divisions of the cells of 
series occurring In general after those of the outer series have taken nlace. 1 
further development consists simply in a continued upward growth around the products 
of the division of the female organ, accompanied by the separation of a terminal 
portion. The latter is then again separated into two portions, the upper of which 
divides again, and so on, until the number of cells characteristic of 
been forme 
pc 
This process may be made somewhat clearer by reference to fi 
n 
k 
