MONOGRAril OF THE LABOULBENIACE-I^:. 233 
a wall-cell or Avitli a parietal cell. The fourth connection could not be distinctly made 
out, and may well have been broken by the crushing which wns resorted to In order to 
separate the cells from one another. The connections of the basal cells with the wall- 
cells, as they are represented in figs. IC and 17, are very readily dcmonstrntcd. It 
will be noticed that in fig. 17 a portion of the posterior basal cell (o'), as well as its 
connection with the wall-cell (/), Is indicated through the anterior basal cell {d)j the 
connections of which with livo wall-cells (c) and (/) are very distinct. The connection 
of the basal cell (y), shown in fig. 16, is not visible in this instance. 
Having considered these special cases and their modifications as far as the}'' arc at 
present known, it may not be isuperfluous, even at the risk of tedious repetition, l^riefly 
to summarize the general development of the perilhecium and of the structures which 
it contains, since it involves matters of such cojisidcrable Importance that a clear 
miderstandinij: of it is essential. 
Summary of the decclopmait of the perUhcchnn and (f the female srxual organs. The 
peritliGcium arises as a lateral^ rarely iis a terminal organ, and consists at an early 
stage of two superposed cells: an upper, which is alone concerned in the formation t)f 
the female organ, and a lower, from which is developed the perithecium proper. The 
upper cell elongates by terminal growth, and is converted into the procarpe through 
the formation primarily of two cross partitions, by wliich it is separated Into a lower 
portion, the carpogenic cell, always a single cell ; a middle portion, the trichophoric 
cell, also always a single cell, and a terminal portion, the trichogyne, which nniy con- 
sist of a single cell, or, through the formation of cross partitions often accompanied 
by copious branching, of very numerous cells. The free extremities only, of the tri- 
chogyne, are receptive, and conjugate with the antherozoids which adhere to them. 
As a result of this union, the trichogyne soon withers and disappears, while the carpo- 
genic cell undergoes a series of divisions. First, by the formation of two transverse 
septa, it is separated into three superposed cells : the upper and lower constituting 
the superior and inferior supporting cells, respectively, which undergo no further 
changes; while the middle cell of the three, known as the ascogonium, divides, by 
more or less oblique partitions, into from two to nine cells, one of which lies at the 
base of the others and is called the secondary inferior supporting coll; while the one, 
two, four, or eight remaining cells are known as the ascogenic cells. Each nscogenic 
cell then begins at once to produce asci, which bud froui it downward outward and 
upward, and soon becomes quite free In the cavity of the perithecium; destroying, as 
a rule, both supporting cells, and eventually the remains of the trichophoric coll, as well 
as the cells of the perithecium proper (parietal and canal cells) which lie Immediately 
about and above It. 
