MONOGRAni OF THE LABOlTLEKNIACK.r:. 373 
^ 
are remarkably variable. Tho horn-like appcMuin-e, which characterizes the perilhecia of tlio 
species first described, does not jTove of goiieric value, since it is absent in fourfunna. The only 
genus which might possibly be mistaken for it, however, is Sphaieromyces : hui<h..pro.senro in fliis 
case of flask-shapeJ anfhoridia on (lie branches of the appendage, not to monf ion ils wcll-dcvclopcd 
pcrithccial stalk-cell, distinguish it without dilhculty. its grcatesl ],ecnlianty Hrs in the character 
and development of the pcrithecium, which diflfcrs from the more tyj>ical forms Jn important 
points; and although I have been unable to obtain young material which illustrates all the early 
stages, it appears in amirahilis to be as follows : The young individual consists of asimjjlc scries 
of superposed cells, the distal of which begin to branch at an early s«agc; and the first indica- 
tion of the formation of the pcrithecium consists in the division by a transverse septum of ono 
of the^cells of this series. Of the two cells resulting from this division, the upper gn»uh ouf- 
ward and upward, its outgrowth becoming a fingor-likc projection (I'latc XXIV, fig. 4, at tbo 
left) that later elongates to form the fihimentous ti-ichogync. This outgrowth, in flic stnge rop- 
resented by the figure, has become divided by two transverse partitions into (luce cells ; llie upprr 
being the young trichogyne, the cell below it the trichoj)horic cell ; while b.^lou (his, not projeeling 
from the axis of the plant, is the smaller carpogenic cell. At tlic right of flic carpogrnic cell a 
vertical septum has divided the remainder of the original uj»per cell into two ].;irt.s, an outer and 
an axilc cell ; but in other species, like C. cojitortun (Plate XXV, fig. 9), Ibis division smms to 
be absent; The lower of the two original cells has, in the mciin time, been divided by a vertical 
septum into two cells, one of wliich (Piute XXIV, fig. 4, x) has also begun to grow outward and 
upward. As far as can be determined, this cell (x) by further growth .-nul division produces 
only one of the series of wall- and canal-cells ; while the cell previously referred to as the axilc 
cell, lying behind the carpogenic cell, gives rise to the three other series. Having been unable 
to separate the cells in this region and ascertain (heir protoplasmic connections with the cells of 
the pcrithecium, it is uncertain whether the above statement is entirely correct in so far as con- 
cerns the number of wall-cells originating from each of the cells mentioned ; but that they give 
rise to the outer and inner series of i)erithccial cells cannot be doubted. The latter grow uj 
around and beyond (he carpogenic and trichophoric cells, and by the successive division of tl 
terminal members form the mnin l)ulk of the pcrithecium. The base of (he friehon-vnc is thus 
left behind, as it were, in the angle between the pcrithecium and the appendage (Plate XXIV, 
While in other genera, 
\\n 
leir 
17) 
as a rule, only one or two divisions of the pcrithccial cells takes place above the insertion of the 
trichogyne, in the present instance the growth beyond it may be almost indefinite ; as in C. ros- 
ti-atus, for example, or CfiUformis, in each of which (Plato XXIV, fig. 15 and 18) the pcrithe- 
cium is extraordinarily developed. Although the general process by which the pcrithecium 
originates is thus very similar to that which has been described as characteristic of Stigma- 
tomyces or Laboulbenia, it will be observed that its basal cells are differentlv arramjed, nnd 
originate in a different manner. Whether it is possible to homologize the typical stalk-cell and 
secondary stalk-cell with any of those present in this case I do nof feel able to determine. Tho 
pcrithccial cells of the inner series differ from those of other genera from the fact that they seem 
to divide quite independently of, and less frequently than, tlic wall-ccILs ; as may be seen in 
fig. 8, Plate XXIV, the septa of the wall-cells {w c) and of the canal-cell {c i) in this instance 
being by no means coincident. As in other cases, thecanal cells are entirtly destroynd bv the 
