76 SIR DIETRICH BRANDIS ON THE 
Nees. Bambusa verticillata, Willd., is a synonym of Gigantochloa verticillata, Munro, 
but there is no Bambusa verticillata, Nees. 
Among the numerous species of which I have examined transverse sections, Jfelocanna 
bambusoides (Pl. 18. fig. 30 and Pl. 14. fig. 31) is the only one which in the intermediate 
cells bas folds from the lower edge only. For purposes of classification I am not 
disposed to attach any importance to this character. 
The cells of the chlorophyll parenchyma are arranged in horizontal lines parallel to 
the surface of the leaf and they are separated by intercellular spaces. Pl. 11. fis. 5 
represents a flat section showing the tissue below the upper epidermis of Arwndinaria 
Hookeriana, Munro, between a longitudinal nerve and the lower portion of the bulliform 
cells, while Pl. 14. fig. 35 represents the tissue separating the apparent cavities, in a flat 
section made near and parallel to the lower epidermis of Jelocanna bambusoides. Tt will 
be noticed that the chlorophyll-cells, which a transverse section shows as oblong plates, 
in these horizontal sections appear oblong with rounded ends with prominent circular 
markings caused by the tubular folds proceeding from the upper and lower edge of these 
cells. As already stated, this remarkable appearance of the chlorophyll-cells of Bamboos 
was clearly explained by Karelstschicoff in 1868. 
MIDRIB, LONGITUDINAL NERVES, TRANSVERSE VEINS, AND STRANDS OF 
SCLERENCHYMATOUS FIBRES. 
The strueture of the longitudinal nerves in Grasses has been fully described by 
previous authors. With very few exceptions the leaves of Bamboos have a distinct 
midrib, though in many species (e. g. Melocanna bambusoides) the midrib gets very thin 
towards the apex and sometimes cannot be distinguished from the lateral nerves. On 
both sides of the midrib there are generally two classes of longitudinal nerves, stout or 
coarse, and between each pair of these a varying number of fine or slender nerves. The 
former Ettingshausen designates as Seitennerven, Gamble as Secondary Nerves, while the 
thin nerves are called Zwischennerven by the former and Intermediate Nerves by 
the latter author. Each nerve consists of two portions, a vascular bundle and two 
girders of sclerenchymatous fibres, one above, the other below the vascular bundle. 
‘The lower girder frequently is extended upwards, so as partially to enclose the vascular 
bundle (Sasa paniculata, Bambusa vulgaris, Dinochloa andamanica). The stout nerves 
always have from two to four large vessels, in the slender nerves the vessels are 
much smaller. 
The vascular bundles in the longitudinal nerves of Bamboos are enclosed in a sheath — 
which consists of two portions. The outer portion (* Parenchym-Scheide, ” Schwendener) 
consists of one layer of large elongated parenchymatous cells with square ends, which 
contain starch and frequently chlorophyll. In a transverse section this sheath appears 
as a ring of orbicular cells, the walls of which, as a rule but not always, show the 
reaction of cellulose. In some eases, chiefly on the underside, where the lower strand of - 
sclerenchymatous fibres is enlarged, this ring is frequently interrupted. The transvers? — 
