ANATOMY OF ENHALUS ACOROIDES. 365 
flower has 3+3 perianth-leaves (sepals and petals) and 3 stamens, all free and in three 
alternating whorls. 
Each stamen consists of a short stout filament bearing two large bilocular anther-lobes, 
each loculus of which contains from 7 to 14 large pollen-grains. 
In the female inflorescence the peduncle is prolonged, and the flower arises just above 
the point of insertion of the bracts. There is one epigynous female flower composing 
the whole inflorescence. It has 34-3 free perianth-leaves and a synearpous unilocular 
inferior ovary with from 6 to 8 carpels. On the outside this is concealed by mucilage- 
producing papillæ of structure similar to those on the bracts. The ovules are anatropous 
and inserted at different levels in different carpels. 
ANATOMY OF THE Bracts. 
The bracts resemble the leaf generally in structure, but clearly show outstanding 
differences in transverse section. The structure varies at different levels, at the apex 
resembling that of the leaf more or less and at the basal region bearing a resemblance to 
that of the peduncle from which it gradually differentiates. 
As in the leaf, there is an epidermal layer, while the space between the dorsal and 
ventral faces is occupied by lacunar tissue. According to the level, there are from one 
to several subepidermal layers of parenchyma. Near the base there are from one to 
several layers on the adaxial and abaxial side respectively; higher there are one or two 
layers both adaxially and abaxially. The lacunz in the basal region are irregular in 
size and shape, the trabecule running obliquely, longitudinally, or horizontally, and 
not all at right angles with respect to the dorsal and ventral surfaces as in the apical 
region of the bract and in the leaf. At irregular intervals transverse diaphragms, one 
cell thick, span the lacunze. 
The vascular bundles run in the ends of the trabecul as in the leaf. There are a 
large median and two “ marginal bundles," which arise as branches of the central median 
bundles of the peduncle. ‘The large median bundle, however, is on the adaxial side of 
the bract, and a small bundle only is on the abaxial side. Between the median and 
marginal bundles there are smaller bundles of two sizes, alternately larger and smaller, 
on the abaxial side of the bract. Near the apex these send anastomosing branches to 
the adaxial side as in the leaf. The structure of all these bundles is the same as that 
of the leaf-bundles. When old those in the female inflorescence show much thickened 
bast-fibres, but in the male very little had arisen in the oldest specimen examined. 
ANATOMY OF THE MALE INFLORESCENCE. 
The Peduncle.—In transverse section the shape is elliptical. grasa iR 
Peduncle there is a well-defined epidermis, outside which there is à x dun ic m 
The epidermal layer consists of regular cells, the outer walls of which are much thicken 
and show definite layering after treatment with Eau de Javelle. p eee 
Within the epidermis there is a. band of continuous parenchyma (text-fig. 8, p 
is -fig. 8,7) which, 
no lacunæ, This layer, 2 to 3 cells thick, surrounds lacunar tissue po eed { 
except for two vascular bundles (text-fig. 8, a, a), fills the centre. sie 
