392 DR. E. A. NEWELL ARBER ON PSYGMOPHYLLUM FROM 
in Newfoundland, and this fact, in addition to their excellent preservation which is a 
rare feature among Devonian plants, favours the conclusion that the beds are of Lower 
Carboniferous age. 
I have also had opportunities of examining some examples of the British repre- 
sentative of the genus, Psygmophyllum flabellatum (Lindl. & Hutt.), a very rare Coal- 
Measure plant, known only from this country and from a single coalfield. As no 
detailed account of this species has been published, beyond the brief diagnoses given by 
Lindley and Hutton, Schimper and Renault, I have added a full description of it here. 
I have also taken the opportunity of revising the genus as a whole, so that the present 
communication is practically a monograph of the subject. 
Finally, I have ventured to conclude with some discussion on the affinities of 
Psygmophyllum, so far as it is possible to arrive at any provisional views on these 
fossils at the present time. 
I desire to express my obligations to those who have kindly lent me specimens of 
P. flabellatum (Lindl. & Hutt.) In particular, E would express my indebtedness to the 
Council of the Sunderland Publie Museum, who, through the kind offices of the Director, 
Mr. J. A. Charlton Deas, F.R.Hist.S., and his assistant, Mr. G. H. Dutton, have lent me 
their valuable series of specimens of this fossil. Iam likewise under obligations to the 
Council of the Hancock Museum, Neweastle-on-Tyne, who, through their Curator, 
Mr. E. L. Gill, M.Sc., have lent me specimens from the Hutton Collection. To my 
friend Professor Nathorst I would return my thanks for the photograph of his type- 
specimen of P. Williamsoni in the Stockholm Museum, which he has kindly allowed 
me to reproduce here. 
2. PSYGMOPHYLLUM MAJUS, sp. nov. 
(PL 42. fig. 1; Pl. 43. figs. 5, 6; Pl. 44. fig. 9.) 
1910. Psygmophyllum sp., Arber, Proc. Camb. Phil. Soc. vol. xv. p. 391. 
Diagnosis.—Leaves flabellate, large, often exceeding 16 cm. in length and 15 cm. in 
breadth, broadest at the apex, suddenly contracted below to a broad, elongate, ? sheathing 
base. Margins curved. Apex rounded, ? entire, or broadly lobed. Veins radiating 
from the base, with frequent dichotomy throughout the lamina, fairly distant, few in 
number below, and erect but not converging at the apex. 
Type-specimens.—Nos. 2210-2215 and 2088 in the Carb. Plant ColL, Sedgwick 
Museum, Cambridge. 
Description of the specimens.—The specimens consist of several fragmentary impres- 
sions of large, detached, fan-shaped leaves, occurring on slabs of sandy shale. One of 
these leaves (No. 2210) which is fairly complete is seen on Pl. 42. fig. 1. Its length is 
13:5 cm. and its greatest breadth 10:3 em. There is considerable doubt whether the 
actual apex is seen in this specimen. I am inclined to think that the rounded, entire, 
upper extremity is not the true apex, for the apical part of the leaf appears to have been 
broken off sharply, owing to the fact that the plane of the impression of the leaf did not 
