126 STRONG. [Vol. X. 



near the surface to a certain extent before terminating (Figs. 

 20 and 21). These terminations are often not simply rounded 

 knobs but have a more elongated club shape and are somewhat 

 irregular in outline (Figs. 18-21). As a number of fibres press 

 in from all sides towards the centre of the depression in the 

 epithelium formed by the gland, this locality is quite filled with 

 these bodies. The irregularity and size of these enlarged ter- 

 minations may be due to some irregularity in the staining, but 

 since they occur often in the cleanest impregnations and since 

 expansions of even greater size exist elsewhere, e.g., in termina- 

 tions in muscles, they may be considered true pictures. 



These appearances in the glands seem to me to be not easily 

 reconcilable with Dogiel's (14) denial of free endings and 

 assertion of the prevalence of a closed network as the terminal 

 apparatus. It is, of course, possible that these enlargements 

 which lie immediately below the surface, almost in it, are not 

 the final terminations and that there are always, e.g., unstained 

 transverse fibres connecting them and forming closed meshes. 

 It is also true that the various methods of staining nerve fibres, 

 especially the Golgi method, are irregular and incomplete in 

 their action, yet it is not likely that the latter would always omit 

 certain fibres such as these hypothetical ones. The manner in 

 v/hich these fibres terminate negatives still more strongly their 

 existence. That true anastomoses may occur is not to be 

 denied and sometimes the appearances favor their existence 

 {vide supra) but they can hardly be of universal occurrence. 

 Neither physiologically nor embryologically would there seem 

 to be any special reason for their existence in such peripheral 

 structures as epithelium, though they might easily occur now 

 and then owing to secondary fusions. 



The R. hyomandibularis facialis, as described above, leaves 

 the Gasserian ganglion at about the same transverse plane as 

 the R. palatinus (760 ±). As it leaves, it is composed, as has 

 been seen above, of three components, occupying the nerve in 

 the following order : The most dorsal is the dorsal VII com- 

 ponent {Yt, VII b) ; next to this is the motor component (VII ah) ; 

 and most ventral is the fasciculus communis component (S[\\aa) 

 (PI. X, Fig. 26). The destination of the first and last compo- 



