1 38 STRONG. [Vol. X. 



and has traced it caudad into the lateral columns of the cord. 

 According to Osborn it contributes, in Cryptobranchus (45), 

 to at least two of the Vagus roots. 



This root is very probably motor, but would appear to be 

 quite different in character from the ventral motor rootlets 

 mentioned above. 



A further description of the fasciculus communis will be 

 found in another place in this paper. 



All these roots now enter the vago-glossopharyngeal gan- 

 glionic complex. The general shape of this complex is indi- 

 cated in the chart. It curves around the posterior end of the 

 auditory capsule, as though pushed back by the extension of 

 the latter, a peculiarity probably due to the very anterior po- 

 sition of the gill clefts and other parts relatively to the auditory 

 capsule. The ganglion attached to its inner portion posteriorly 

 is the sympathetic ganglion. It is drawn too large in the 

 chart. 



It is not possible to trace the exact relations of the various 

 roots through this complex ; yet, the composition of the latter 

 can, I think, be determined in a general way, and sufficiently 

 to throw considerable light upon its morphology. 



In the first place, it appears that this complex falls into three 

 main divisions which are indicated by the shading in the chart. 

 The most proximal ganglionic division is connected with the 

 nerves before they pass around the hinder apex of the auditory 

 capsule. Immediately beyond this lies another ganglion, and 

 still further along, and quite upon the outside of the auditory 

 capsule, the third ganglion. It will be necessary to describe 

 the various branches of the IX + X before discussing the char- 

 acter of these three ganglia. 



The first root, on account of its large fibres, can be readily 

 followed, and its destination may be finally treated here. Just 

 before entering the vago-glossopharyngeal ganglionic complex 

 it divides. The dorsal division (/), remaining in the dorsal 

 part of the ganglion, curves around the auditory capsule and 

 separates. The ventral division (5) becomes separated from 

 the dorsal by an intervening bundle of fibres. It remains 

 longer in connection with other fibres of the IX -|- X, pro- 



