112 THAXTKK. — MONOGRAPH OF THE LABOULBENIACE.E. 
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plane, and other branches coming from cell (h) both above and below, are not shown. The parietal 
brandies appear to be developed near the bases of the wall-cell branches, but are small and hardly dis- 
tinguishable, their functions being usurped as will appear later, by the wall-cells themselves. 
K\< n while this development of branches is taking place the two basal cells (o) and (h) begin to be 
pressed down by the structures to which they have given origin above, into the cell below from which 
thev originated, and as the development of the ascogonium proceeds this cell, like the cells above it, loses 
its identity ;h ;i cell; the whole series being transformed to a continuous cavity within which the whole 
nseogenie apparatus eventually lies quite free. The procarp resembles other procarps of this group, 
consisting of an ascogonium, an inferior and a superior supporting cell, followed by a trichophoric cell; 
above which the unicellular, sparingly branched or tabulated trichogyne emerges, as has been already 
lescribed (figs* 1 1 12). As the perithecium matures and the ascogenic cells become active, only two or 
thn -e wall-cells which have become indurated and cohere closely to the inner surfaces of the cells within 
which they were formed, persist, as shown in the bent termination of fig. 7, to form the tip of the functional 
perithecium. The cavity of the perithecium is thus the cavity of the cell from which the perithecial branch 
system originated combined with the cavities of the cells through which these branches penetrated; and 
its walls, except at the tip, as above mentioned, are the walls of these cells. The structure may thus be 
termed ;i fixe udo per it hecvum. 
If then one examines the conditions found when the plant is fully mature it will be seen to consist 
of the same three regions distinguished in the young condition. The terminal region, however, above 
the appendiculate cells, excepting only its basal cell which acts as a false stalk-cell to the pseudoperithe- 
eiimi, has become transformed into a single chamber, the wall of which corresponds to the combined 
walls of the four or more cells which were superposed above the basal cell of the series comprising 
this distal region. Within this chamber have developed from the subbasal cell of the series, structures 
which correspond to all those found in the normal perithecium of the Laboulbeniales. Of these struc- 
tures, however, nil are eventually disorganized with the exception of a few small wall-cells, which persist 
distally and are functional in regulating the spore-discharge, and the ascogenic cells, of which there appear 
to be four. These ascogenic cells, then, together with the usual mass of asci and spores, float free within 
a structure resembling a perithecium and performing the same function, but which when viewed super- 
ficially has absolutely nothing in common with any of the structures seen in perithecia of the normal type. 
Such an endogenous origin of the perithecium has no parallel in the other genera, unless it be in 
Zodiomyce* Mid perhaps also in Euzodivmyces. In the former the perithecigerous area arises endoge- 
nously in a somewhat similar fashion, and if in the present instance the perithecial structures persisted, 
and the walls of the cells into which they penetrate were eventually destroyed, the conditions would be 
closely comparable: the differences depending merely on the differences in the general cell structure of 
tl 
ie 
many 
pe 
B body of the plant in either ease; the complicated multicellular body of Zodiomyces producing mar 
rit In . i;i where the present simple type gives rise to but one. The character and origin of the antheridial 
branches in the two cases are however totally different. 
The antheridial appendages in the present genus are peculiar, from their position on the primary axis 
below the perithecinm, a condition seen nowhere else unless possibly in C hcetomuees ; although such an 
trrangement, as in Chmatomyces, is often seen in branches from this axis. The antheridial cells them- 
selves are merely the lower cells of the appendage or its branches (fig. 13), from which short outgrowths 
arise that are converted into discharge-tubes through which the sperm-cells make their exit as in simple 
antheruha of the ordmary type. The conditions present in the antheridial appendages in this instance do 
not therefore appear to differ essentially from those seen, for example, in Rhadinomyces; except that the 
enes of antheridial cells is not definitely differentiated. At the same time the conditions described are 
very similar to those m Rhynchophoromyces and illustrated in my Monograph, Plate XXIV, fig. 24: but 
a hether the sperm-cells , n the present instance possess a wall, as in Rhynchophoromyces, I am not able to 
— . 1 he ant her.dial characters of Coreomyces thus tend to break down the distinction formerly assumed 
sa v 
