Notes on Hawaiian Zonitidae and Succineidae 
By C. MontacuE Cooke, Jr. 
INTRODUCTION 
The material on which the present paper is based forms part of 
the malacological collection of the Bishop Museum. In 1920 I 
took to Philadelphia for comparative study the animals and shells 
of Hawaiian representatives of the families of Zonitidae, Endodon- 
tidae, and Succineidae and worked out there the anatomy of a 
number of the Zonitidae and Succineidae, but was unable to com- 
plete the analysis for lack of sufficient material. These notes deal 
with several genera of Succineidae and with three of the rarest 
genera of Zonitidae found in Hawaii. For comparison with the 
Hawaiian forms the very rich collection of wet material in the 
Academy of Natural Sciences was placed at my disposal in Phila- 
delphia by Dr. Henry A. Pilsbry, to whom I am also indebted for 
advice and many courtesies. Thanks are due also to Miss Helen 
Winchester, who made the drawings for the two plates. 
ZONITIDAE 
GODWINIINAE 
Sykes! proposed the generic title of Godwinia for Gould’s 
Vitrina caperata, basing his conclusions on the anatomical studies 
of Godwin-Austen. From a further study of the anatomy it appears 
that a new subfamily title is necessary. The shells are rather small, 
8-13 mm. in diameter, of few rapidly enlarging whorls, and a large 
aperture; the embryonic whorls are more or less distinctly and 
roughly, distantly and radiately costate; the umbilicus is small and 
circular. 
Unfortunately the animals in all the specimens examined were 
much contracted. As shown by Godwin-Austen there are no shell- 
lobes. Both right and left dorsal lobes are strongly developed. 
The sexual orifice is situated rather high and is back of the pul- 
monary orifice (fig. 3, @). There is no tail pore or slit. The penis 
is entirely different from that of other zonitoid snails. It is rather 
large with a very short stout retractor. The distal end is slightly 
enlarged into a head and is very thick walled with a minute almost 
* Sykes, E. R., Mollusca: Fauna Hawaiiensis, ii, p. 277, 1900. 
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