WILLIAMS: THE GENUS DESMATODON 211 
TYPE LOCALITY: Rocky Mountains of British America. 
DistRIBUTION: Greenland; Beechey Island, Arctic America; the 
Canadian Rockies; also in Europe. 
ExsfccATAE: Drummond, Musci Am. 145. 
ILLUSTRATION: B.S.G. Bryol. Eur. pl. 136. 
3. DESMATODON GUEPINI B.S.G. Bryol. Eur. (18-20): Desma- 
todon 8. 1843 
Trichostomum Guepini C. Miill. Syn. 1: 590. 1849. 
Barbula Guepini Schimp. Syn. ed. 2, 197. 1876. 
Tortula Guepini Broth. in E. & P. Nat. Pfl. 1°: 430. 1902. 
_ Autoicous, the two or three very small male flowers scattered 
along the stem and composed of four or five pale, ecostate, ovate, 
acute leaves, smooth or nearly so, the outer longer ones about 0.5 
mm. long, enclosing three or four antheridia about 0.25 mm. long, 
with few or no paraphyses: fertile plants rather loosely cespitose, 
bud-like, 1-3 mm. high; the larger upper leaves ovate to somewhat 
spatulate, with blade 1-1.5 mm. long, the apex somewhat rounded 
or acute, the margins entire and revolute from near the apex al- 
most to base and the costa mostly smooth on the back, excurrent 
into a nearly smooth point one fifth to one half the length of the 
blade; costa in cross-section showing mostly two guide-cells, four 
or five cells of about the same size in one row on the ventral side 
and on the dorsal side a thick stereid band with outer cells differ- 
entiated; leaf-cells rather obscure and densely papillose in upper 
part of leaf, more or less four to six sided, not or scarcely elongate, 
14-16 u wide, those of basal part smooth, pale, larger, more or less 
rectangular: outer perichaetial leaves not differentiated, the inner 
small, acutely pointed, with flat margins; seta erect, about 8 mm. 
long; capsule erect, oblong-cylindric, 1-1.5 mm. long without 
lid, the stomata in one row near the base; annulus narrow, per- 
sistent, of one or two rows of cells; peristome pale, densely papil- 
lose, of sixteen slightly oblique, narrow teeth mostly divided nearly 
to the base into two filiform forks from a basilar membrane ex- 
tending well above the annulus; lid high-conic, its height about 
twice the basal diameter, the cells a little above the base elongate 
in nearly erect rows; spores nearly smooth, the larger 164 in diam- 
eter; calyptra cucullate, descending about half way down the cap- 
sule. [FiIG. 3. 
TYPE LOCALITY: France. 
DistTRIBUTION: California and France. 
ILLUsTRATION: B.S.G. Bryol. Eur. pl. 133. 
