236 GLEASON: TAXONOMIC STUDIES IN VERNONIA 
Minnesota, the Ohio State University,and the University of Wis- 
consin for the loan of material; to the directors or curators of the 
Brooklyn Botanic Garden, the Gray Herbarium, the National 
Herbarium, the New England Botanical Club, and the Phila- 
delphia Academy of Sciences for the facilities of their herbaria and 
libraries; to Dr. B. L. Robinson, Dr. J. M. Greenman, Mr. W. 
R. Maxon, and Mr. E. E. Watson for information, assistance, and 
criticism; and especially to the University of Michigan, where 
part of the work was done. 
VERNONIA BORINQUENSIS Urban 
In describing the species, Urban took as the type a form with 
hirsute achenes, resinous-dotted leaves, and short straight ap- 
pressed hairs on the lower leaf surface. This is the commonest 
form in American herbaria. Urban’s variety Stahlii covers a 
much rarer form with glabrous achenes and without resinous dots 
on the leaves. ' Two other forms also occur, which are deserving of 
varietal names. 
Vernonia borinquensis resinosa var. nov. 
Achenes glabrous; leaves conspicuously dotted with glands and 
resin on the lower surface: otherwise like the typical form of the 
species. 
TYPE: ex herb. E. W. D. Holway, collected at Cayey, Porto 
Rico, January, 1911, and deposited in the herbarium of the New 
York Botanical Garden. 
Vernonia borinquensis hirsuta var. nov. 
_ Achenes hirsute; leaves densely sericeous-hirsute beneath 
with bent or curved hairs 2-4 mm. long, nearly or completely 
concealing the resinous dots; otherwise like the typical form of the 
species. 
Type: Britton, Stevens, & Hess 2471, collected in a wooded 
valley, Rio de Maricao, Porto Rico, at an altitude of 500-600 
meters, April 2, 1913, and deposited in the herbarium of the New 
York Botanical Garden. Field notes indicate that it is a vine, 
two meters long. 
The species and its three varieties may be distinguished as 
follows: 
