44 
P. blanda “werden bei beiden Schritten der Meiosis Wande 
gebildet von sehr unregelmiisziger Lage, die jedoch nach kurzer 
Zeit wieder aufgelést werden.* In this case also we can safely 
accept the formula AAAA—TIIIa—3ab. 
Arno.pi’s publication on the Huphorbiaceae is most annoying inits 
lack of detail. Moreover there are absolutely no plates and the pu- 
blished figures are too few in number to base any conclusions 
upon. The author seems to have suffered from lack of material. 
Ceramanthus’ mature embryosac probably contains only the 
four nuclei of the micropylar end*..... am entgegengesetzten 
Ende sah ich keine Kerne, obgleich dies nicht fir absolut 
gewiss gelten kann.‘ The only figure of the two-nucleate stage 
shows clearly a primary chalazal nucleus as well as the pri- 
mary micropylar one. The next figure represents four nuclei 
at the micropylar end. Evidently the chalazal nucleus degene- 
rates very soon after its formation, which is expressed by the 
formula A—I—4z, The life history of the Codiaeum is confined 
to five lines and four figures. There seems to be a four-nucleate 
sac at the end, but how it originated cannot be decided. Ar- 
NoLpI’s figures point to some development of the primary cha- 
lazal nucleus. Possibly the sac-nucleus in full-grown state is 
the result of a fusion of two polars, the sac thus being not 
four- but actually five-nucleate, and the formula A—I—3az. 
Pedilanthus as a rule corresponds to A—I—4. Occasionally the 
reduction has not gone so far, there being two antipodals 
left: A—T— 2a. 
The 16-nucleate Euphorbiaceae, described by Movitewski (1909, 
1910, 1911) and by Arnoxpr (1912) are of such remarkable uni- 
formity as to the development of their embryosacs that it is 
not necessary to treat them separately. Vacuolation does not 
commence before the eight-nucleate stage, which proves a tetra- 
sporic condition. Each of the four megaspores develops only a 
“micropylar group*; the chalazal groups are wholly suppressed 
and not even a primary chalazal nucleus appears. The four 
“micropylar groups“ are to be found: one at the micropylar 
end of the sac, one at the chalazal end and one at each end 
of a transverse axis. Each group organizes an egg-apparatus, 
