5 
lated and shortly afterwards is found divided by cell walls. 
(Plate VI, Fig. 32). At this stage seeds have already attained 
a length of about 8 mm. and the fruits even of 15 to 25 cM. 
Summary. 
The archesporium of Moringa oleifera Lam. consists of one 
single cell. 
Four megaspores are formed, the outer two lying rectangular 
to the axis of the sporangium. 
The inner megaspore is the functioning one. By two suc- 
cesive divisions the normal polarised four-nucleate stage is rea- 
ched. The third division is restricted to one of the micropylar 
nuclei only, the other micropylar and both chalazal nuclei 
remaining undivided. 
Thus the mature embryosac is five-nucleate, showing a nor- 
mal egg apparatus of two synergids (sisters) and the egg. The 
position of the embryosac nucleus is taken by the two chalazal 
nuclei, lying in close contact and quite near the egg. 
_ At fertilization one male nucleus fuses with the eggnucleus 
and the other male nucleus enters in triple fusion with the 
chalazal nuclei. This primary endosperm nucleus at once com- 
mences to divide and soon numerous free endosperm nuclei are 
present. The egg remains undivided for quite a long time. 
Embryo formation begins with free nuclear division up to the 
sixteen-nucleate stage. Then cell walls are formed and further 
divisions no longer take place simultaneously. The embryo 
rapidly grows and the cytoplasm in which the endosperm nuclei 
are embedded is divided by cell walls. 
This way of megaspore — and of embryosac formation, lea- 
ding to a 5-nucleate sac of a very special character, up till 
now has only been reported for Garcinia (M. Taevs: ae du 
Jard. Bot. de Buitenzorg 1910: Le sac embryonnaire et Vem- 
bryon dans les Angiospermes. Nouvelle série de recherches, I 
Garcinia Kydia (Roxs.), Garcinia Treubii (Pierre) ). 
