Coussarea Aubl., Hist. PI. Guiane 1: 98. 1775. 

 Type: Coussarea violacea Aubl. 



Coussarea austin-smithii Standi., Publ. Field 

 Mus. Nat. Hist., Bot. Ser. 18: 1286. 1938. 

 Type: COSTA RICA: Region of Zarcero, 

 4700 ft., 25 Aug 1937, A. Smith 4226 

 (Holotype F 905233 (photos MEXU, PTBG)). 



Coussarea caroliana Standi., Publ. Field Mus. 

 Nat. Hist., Bot. Ser. 22: 178. 1940. Type: 

 COSTA RICA: Alajuela, Canton San Carlos, 

 Suerre, 1000 m, 2 Mar 1939, A. Smith H1645 

 (Holotype F 981032 (photo PTBG); Isotype 

 US (photos MEXU, PTBG)). 



Coussarea cerroazulensis Dwyer & Hayden, 

 Ann. Missouri Bot. Gard. 53: 374. 1966. 

 Type: PANAMA: Panama, Cerro Azul, 600 

 m, 6 Aug 1961, J. D. Dwyer 1368 (Holotype 

 MO 1810161 (photos MEXU, PTBG)). 



Coussarea chiriquiensis (Dwyer) C. M. Taylor 

 in W. C. Burger & C. M. Taylor, Fieldiana, 

 Bot., n.s. 33: 113. 1993. Rudgea chiriquiensis 

 Dwyer, Ann. Missouri Bot. Gard. 67: 476. 

 1980. Type: PANAMA: Chiriqui, Cerro 

 Colorado, along road above San Felix, 29 km 

 above bridge over Rio San Felix (7.9 km 

 above tumoff to Escopeta), 1500 m, 14 Jul 

 1976, T. B. Croat 37071 (Holotype MO 

 2798805 (photo PTBG); Isotype MO). 

 — Psychotria nebulosa (Dwyer) C. M. Taylor 



Coussarea colonis Standi., Publ. Field Mus. Nat. 

 Hist., Bot. Ser. 22: 179. 1940. Type: 

 PANAMA: Bocas del Toro, Buena Vista, 

 region of Almirante, Jan-Mar 1928, G. P. 

 Cooper 228 (Holotype F 579282 (photo 

 PTBG)). Norn, superfl. for Coussarea 

 euryphylla Standi., 1929 — Raritebe 

 palicoureoides Wernham subsp. 

 ". H. Kirkbr. 





Coussarea curvigemma Dwyer, Phytologia 38- 

 215. 1978. Type: PANAMA: Barro Colorado 

 Island, Wheeler Trail 850, 1 Jun 1971, T. B. 

 Croat 14863 (Holotype MO 2050013 (photo 

 PTBG)). 



Coussarea dariensis Steyerm., Ceiba 3: 20. 

 1952. Type: PANAMA: Darien, Rio Chico, 



vicinity of Yaviza, 100 ft., 9-1 1 Jun 1947, 

 P. H. Allen 4576 (Holotype F 1397594 (photo 

 PTBG); Isotypes MO, NY, n.v.).— Coussarea 

 paniculata (Vahl) Standi. 



Coussarea durifolia Dwyer, Ann. Missouri 

 Bot. Gard. 67: 124. 1980. Type: PANAMA: 

 Panama, along El Llano-Carti Tupile road, 12 

 miles above Panamerican Highway, 200-500 

 m, 26-27 Mar 1973, R. Liesner 1180 

 (Holotype MO 2219526 (photo PTBG)). 



Coussarea enneantha Standi., J. Wash. Acad. 

 Sci. 18: 282. 1928. Type: PANAMA: Darien, 

 Cana and vicinity, 2000-6500 ft., 17 Apr-8 

 Jun 1908, R. S. Williams 841 (Holotype US 

 678317 (photos MEXU, PTBG)). 



Coussarea euryphylla Standi., Publ. Field Mus. 

 Nat. Hist., Bot. Ser. 4: 294. 1929. Type: 

 PANAMA: Bocas del Toro, Buena Vista 

 camp on Chiriqui trail, region of Almirante, 

 Jan-Mar 1928, G. P. Cooper 228 (Holotype F 

 579282). Dukea, Raritebe.— Raritebe 

 palicoureoides Wernham subsp. 

 dwyerianum J. H. Kirkbr. 



Coussarea garciae Standi., Publ. Field Mus. 

 Nat. Hist., Bot. Ser. 22: 180. 1940. Type: 

 COLOMBIA: Choco, Bahia Solano, along 

 Quebrada Jellita, 50-100 m, 22 Feb 1939, 

 E. P. Killip & H. Garcia 33524 (Holotype F 

 999258 (photo PTBG); Isotype US (photo 

 PTBG)). 



Coussarea hondensis (Standi.) C. M. Taylor & 

 W. C. Burger, Selbyana 12: 138. 1991. 

 Psychotria hondensis Standi., J. Wash. Acad. 

 Sci. 18: 183. 1928. Type: COSTA RICA: Bois 

 de Rio Hondo, versant atlantique, 100 m, Aug 

 1901,//. Pittier 16161 (Holotype US 764151 

 (photos MEXU, PTBG)). 



Coussarea imitans L. O. Williams, Phytologia 

 26: 488, fig. 1973. Type: GUATEMALA: 

 Izabal, along trail beginning from mile 33.23 

 between Dartmouth and Moralaes toward 

 Lake Izabal, Montana de Mico, 30-150 m, 7 

 Apr 1940, J. A. Steyermark 39013 (Holotype 

 F 1042963 (photos MEXU, PTBG)). 



Coussarea impetiolaris Donn. Sm., Bot. Gaz. 

 (Crawfordsville) 37: 418. 1904. COSTA 

 RICA: La Colombiana, Llanuras de Santa 

 Clara, 200 m, Jun 1899, H. Pittier 7582 



