A NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE 



(Holotype US 943498 (photos MEXU, 

 PTBG); Isotype US 943497 (photos MEXU, 

 PTBG)). 



Coussarea jefensis Dwyer, Ann. Missouri Bot. 

 Gard. 67: 128. 1980. Type: PANAMA: 

 Panama, Cerro Jefe, ca. 1000 m, 25 Aug 

 1975, S. Mori & L. Mi 7927 (Holotype MO 

 2285634 (photo PTBG)). 



Coussarea latifolia Standi., J. Wash. Acad. Sci. 

 18: 281. 1928. Type: COSTA RICA: Forets 

 de Tsaki, Talamanca, 200 m, Apr 1895, A. 

 Tonduz 9574 (Holotype US 938840 (photos 

 MEXU, PTBG)). 



Coussarea Ioftonii (Dwyer & Hayden) Dwyer, 

 Ann. Missouri Bot. Gard. 67: 129. 1980. 

 Faramea Ioftonii Dwyer & Hayden, 

 Phytologia 15: 56. 1967. Type: PANAMA: 

 Panama, Cerro Jefe in Clusia forest at 

 2700-3000 ft., 27 Jan 1966, E. L. Tyson, J. 

 Dwyer & K. Blum 3323 (Holotype MO 

 1824505 (photo PTBG); Isotype MO (photo 

 PTBG)). 



Coussarea mediocris Standi. & Steyerm., Publ. 

 Field Mus. Nat. Hist., Bot. Ser. 23: 248. 1947. 

 Type: GUATEMALA: Huehuetenango, Sierra 

 de los Cuchumatanes, vicinity of Maxbal, 

 about 17 miles north of Barillas, 1500 m, 

 15-26 Jul 1942, J. A. Steyermark 48732 

 (Holotype F 1 134264 (photos MEXU, 

 PTBG); Isotype US (photos MEXU, PTBG)). 



Coussarea mexicana Standi., Publ. Field Mus. 

 Nat. Hist, Bot. Ser. 17: 274. 1937. Type: 

 MEXICO: Oaxaca, Ubero, 30-60 m, Jun 

 1937, L. Williams 9442 (Holotype F 897721 

 (photos MEXU, PTBG); Isotypes P (photos 

 MEXU, PTBG), US (photos MEXU, PTBG)). 



Coussarea morii Dwyer, Ann. Missouri Bot. 

 Gard. 67: 130. 1980. Type: PANAMA: 

 Darien, Rio Tuquesa, at middle Tuquesa 

 Mining Company Camp called Charco Peje, 

 tropical wet forest, 250 m, 8 Jul 1975, S. Mori 

 7058 (Holotype MO 2821790 (photo PTBG)). 



Coussarea nebulosa Dwyer, Ann. Missouri Bot. 

 Gard. 67: 131. 1980. Type: PANAMA: 

 Chiriqui, uncut cloud forest at Monte Rey 

 near Boquete, 20 Jul 1971, T. B. Croat 15868 

 (Holotype MO 2162995 (photo PTBG); 



Isotype MO 4043108 (photo PTBG)). 



— Psychotria nebulosa (Dwyer) C. M. Taylor 



Coussarea neei Dwyer, Ann. Missouri Bot. 

 Gard. 67: 132. 1980. Type: PANAMA: 

 Veraguas, 6-7 km W of Santa Fe on new road 

 past agriculture school, 2900 ft., 17 Feb 1974, 

 M. Nee 9820 (Holotype MO 2241791 (photo 

 PTBG); Isotype MO 2241792 (photo PTBG)). 



Coussarea nigrescens C. M. Taylor & Hammel, 

 Selbyana 12: 135, fig. 1. 1991. Type: COSTA 

 RICA: Heredia, Finca La Selva, the OTS field 

 station near Puerto Viejo de Sarapiqui, near 

 the junction of the Rios Puerto Viejo and 

 Sarapiqui, 100 m, 5 Nov 1980, B. Hammel 

 10383 (Holotype DUKE, n.v.). 



Coussarea paniculata Standi., J. Wash. Acad. 

 Sci. 18: 282. 1928, based on Froelichia 

 paniculata Vahl, Eclog. Amer. 1: 3. 1796. Not 

 Froelichia paniculata Moench., 1794. Type: 

 TRINIDAD: Ryan s.n. (C, n.v.). 



Coussarea psychotrioides C. M. Taylor & 

 Hammel, Selbyana 12: 135, fig. 2. 1991. 

 Type: COSTA RICA: Heredia, Finca La 

 Selva, the OTS field station near Puerto Viejo 

 de Sarapiqui, near the junction of the Rios 

 Puerto Viejo and Sarapiqui, 100 m, 12 Jul 

 1979, J. H. Beach 1467 (Holotype DUKE, 

 n.v.; Isotypes CR, n.v., F, MO). 



Coussarea roseocremea Dwyer, Ann. Missouri 

 Bot. Gard. 67: 133. 1980. Type: PANAMA: 

 Panama, El Llano-Carti Highway, 6-10 km 

 north of El Llano, 13 Apr 1973, R. L. Dressier 

 4334 (Holotype MO 2126258 (photo PTBG)). 

 —Psychotria roseocrema (Dwyer) C. M. 

 Taylor 



Coussarea talamancana Standi., Publ. Field 

 Mus. Nat. Hist., Bot. Ser. 18: 1288. 1938. 

 Type: COSTA RICA: Talamanca Valley, Jun 

 1927, G. P. Cooper T4 (Holotype F 572698 

 (photo PTBG)). 



Coussarea veraguensis Dwyer, Ann. Missouri 

 Bot. Gard. 67: 134. 1980. Type: PANAMA: 

 Veraguas, NW of Santa Fe, 1 km from 

 Escuela Agricola Alto de Piedra, on the slopes 

 of Cerro Tute, S. Mori & J. Kallunki 4800 

 (Holotype MO 2276605; Isotype MO 

 2276606).— Coussarea caroliana Standi. 



