i NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE 



Hedyotis sharpii (Terrell) G. L. Nesom, Syst. 

 Bot. 13: 434. 1988. Houstonia sharpii Terrell, 

 Brittonia 32: 490,/?g. 1. 1980. Type: 

 MEXICO: Hidalgo, camino entre Metepec y 

 Tenango de Doria, entre 1 6 y 3 1 .5 km al NNE 

 de Tulancingo, 18 km NNE de Metepec, 29 

 Mar 1974, S. D. Koch 7423 (Holotype US 

 2754863 (photos MEXU, PTBG); Isotypes 

 CHAPA, ENCB, MEXU). 



Hedyotis sinaloeae W. H. Lewis, Rhodora 63: 

 222. 1961, nom. nov. for Houstonia parvula 

 Brandegee, Zoe 5: 221. 1905. Not Hedyotis 

 parvula (A. Gray) Fosb., 1943. Type: 

 MEXICO: Sinaloa, vicinity of Culiacan, 12 

 Oct 1904, T. S Brandegee s.n. (Holotype UC, 

 n.v.; Isotypes GH, n.v., US (photos MEXU, 

 PTBG)). 



Hedyotis spellenbergii G. L. Nesom & 

 Vorobik, Syst. Bot. 13: 432. 1988. Type: 

 MEXICO: Chihuahua, Mun. Ocampo, area of 

 Cascada de Basaseachic at the confluence of 

 Rio Basaseachic and Rio Durazno, ca. 2 mi S 

 of village of Basaseachic, 2100-2200 m, 18 

 May 1985, J. Grimes, T. Avers, M. Lavan & 

 R. Scott 2855 (Holotype TEX, n.v.; Isotypes 

 ARIZ, n.v., CAS, n.v., MEXU, n.v., NMC, 

 n.v., NY, n.v., US, n.v.). 



Hedyotis spinescens Sesse & Moc., Fl. Mexic. 

 22. 1893. Type: MEXICO: Habitat in aridis 

 Tehuacani circuitibus. Floret Maio, Sesse & 

 Mocino s.n. (presumably MA, n.v.). 

 — Bouvardia erecta (DC.) Standi. 



Hedyotis teretifolia (Terrell) G. L. Nesom, 

 Syst. Bot. 13: 434. 1988. Houstonia teretifolia 

 Terrell, Brittonia 31: 166. 1979. Type: 

 MEXICO: Coahuila, 12 km NNE of Las 

 Margaritas on easternmost ridge of Sierra de 

 las Margaritas, 1300-1400 m, 24 Sep 1972, 

 M C. Johnston, F. Chiang & T. Wendt 9508E 

 (Holotype TEX, n.v.). 



Hedyotis terrellii Lorence, Novon 7: 51, fig. 3. 

 1997. Type: MEXICO: Oaxaca, Distr. de 

 Mixe, Mun. de Totontepec, 3 km al SW de 

 Totontepec, carretera a Mitla, 1910 m, 6 Sep 

 1986, R. Torres C. & C. Martinez 9067 

 (Holotype PTBG 12314; Isotype MEXU). 



Hedyotis umbratilis (B. L. Rob.) W. H. Lewis, 

 Rhodora 63: 222. 1961. Houstonia umbratilis 



B. L. Rob., Proc. Amer. Acad. Arts 45: 401. 

 1910. Type: MEXICO: Nuevo Leon, shaded 

 cliffs and mountains, near Monterrey, 25 Apr 

 1906, C. G. Pringle 13877 (Holotype GH, 

 n.v.; Isotypes MO (photos MEXU, PTBG), 

 US (photos MEXU, PTBG)). 



Hedyotis vergrandis W. H. Lewis, Rhodora 63: 

 222. 1961. Nom. nov. for Houstonia prostrata 

 Brandegee, Zoe 5: 105. 1901. Not Hedyotis 

 prostrata Blume, 1823, nor Korth., 1851. 

 Type: MEXICO: Baja California, La Palma, 

 Cape Region, 25 sep 1899, T. S. Brandegee 

 s.n. (Holotype UC, n.v.; Isotypes GH, n.v., 

 US, 2 sheets (photos MEXU, PTBG)). 



Hedyotis watsonii W. H. Lewis, Rhodora 63: 

 222. 1961. Nom. nov. for Oldenlandia ovata 

 S. Watson, Proc. Amer. Acad. 18: 97. 1883. 

 Not Hedyotis ovata Thunb. ex Maxim., 1833, 

 nor Thwaites ex Hook, f., 1892. Type: 

 MEXICO: Nuevo Leon, Guajuco, Feb-Oct 

 1880, E. Palmer 399 (Lecototype GH, n.v.; 

 Isolectotype US 47919 (photos MEXU, 

 PTBG)). 



Hedyotis wrightii (A. Gray) Fosberg, Lloydia 

 4: 290. 1941. Houstonia wrightii A. Gray, 

 Proc. Amer. Acad. Arts 17: 202. 1882. Type: 

 MEXICO: Nuevo Leon, shaded cliffs of 

 mountains, near Monterrey, 25 Apr 1906, 



C. G. Pringle 13877 (Holotype GH, n.v.; 

 Isotypes MO (photos MEXU, PTBG), US 

 (photos MEXU, PTBG)). 



Hedyotis xestosperma (B. L. Rob. & Greenman) 

 W. H. Lewis, Rhodora 63: 222. 1961. 

 Oldenlandia xestosperma B. L. Rob. & 

 Greenman, Proc. Amer. Acad. Arts 32: 41. 

 1896. Type: MEXICO: Oaxaca, open glades, 

 Sierra de San Felipe, 8000 ft., Jun 1894, C. G. 

 Pringle 4692 (Lectotype GH, n.v.; 

 Isolectotypes BM, n.v., F, K, n.v., MO 

 (photos MEXU, PTBG), MSC, n.v.. PH. n.v.. 

 US, 2 sheets (photos MEXU, PTBG), VT 

 n.v.); designated by Terrell, 1980). Houstonia. 

 — Bouvardia xestosperma (B. L. Rob. & 

 Greenman) Terrell & S. D. Koch 



