(Holotype US 1315865 (photos MEXU, 

 PTBG); Isotype F).— Hedyotis drymarioides 

 (Standi.) W. H. Lewis 



Houstonia fasciculata A. Gray, Proc. Amer. 

 Acad. Arts 17: 203. 1882. Syntypes: U.S.A.: 

 Texas, Presidio, Bigelows.n. (GH); New 

 Mexico, Organ Mountains, G. R. Vasey s.n. 

 (GH, n.v.); MEXICO: Coahuila, near Parras 

 and Monclova, E. Palmer 404 (GH, n.v.), E. 

 Palmer 406 (GH, n.v.).— Hedyotis intricata 

 Fosberg 



Houstonia fruticosa Willd. ex Roem. & Schult., 

 Syst. 3: 527. 1818. Type: VENEZUELA: 

 Monagas, Caripe, Humboldt s.n. (B, 

 presumably destroyed).— Declieuxia 

 fruticosa (Willd. ex Roem. & Schult.) Kuntze 



Houstonia fruticosa Rose, Contr. U.S. Natl. 

 Herb. 1: 132. 1892. Not Hedyotis fruticosa L, 

 1753, nor Willd. ex Roem. & Schult., 1818. 

 Type: MEXICO: Baja California, Carmen 

 Island (Isla Carmen), 1-7 Nov 1890, E. 

 Palmer 885 (Holotype US 47256 (photos 

 MEXU, PTBG); Isotypes MO (photos 

 MEXU, PTBG), US (photos MEXU, PTBG)). 

 — Hedyotis mucronata Benth. 



Houstonia gracilenta I. M. Johnst., Proc. Calif. 

 Acad. IV, 12: 1174. 1924. Type: MEXICO: 

 Baja California, San Diego Island, rocky 

 island slope facing sea, 2 May 1921, 1. M. 

 Johnston 3927 (Holotype UC, n.v.; Isotypes 

 GH, n.v., US (photos MEXU, PTBG)). 

 —Hedyotis gracilenta (I. M. Johnst.) W. H. 



Houstonia gracilis Brandegee, Zoe 5: 238. 

 1907. Not Hedyotis gracilis DC, 1830. Type: 

 MEXICO: Veracruz, Orizaba, May 1904, 

 C. A. Purpus 1251 (Holotype UC, n.v.; 

 Isotypes F, MO (photos MEXU, PTBG)). 

 —Hedyotis exigula W. H. Lewis 



Houstonia humifusa (A. Gray) A. Gray, Proc. 

 Amer. Acad. Arts 4: 314. 1859. Hedyotis 

 humifusa A. Gray, Boston J. Nat. Hist 6" 216 

 1850. Type: U.S.A.: Texas, near 

 Fredrickburg, 1847, F. Lindheimer 62 1 

 (Holotype GH, n.v.; Isotypes BM, n.v., K, 

 n.v., MICH, n.v., MO, PH, n.v., US). 



2 kingii Terrell, Brittonia 32: 491, fig. 

 . Type: MEXICO: Oaxaca, E-facing 



slopes, mountains along route 175, 28 km by 

 road south of Valle Nacional on way to Ixtlan 

 de Juarez, ca. 2000 m, 30 Jul 1959, R. M. 

 King 2107 (Holotype MICH, n.v.; Isotype 

 MICH, n.v.).— Hedyotis kingii (Terrell) G. L. 

 Nesom 



Houstonia latifolia Willd. ex Roem. & Schult., 

 Syst. 527. 1808. Not Hedyotis latifolia (M. 

 Martens & Galeotti) Walp., 1846. Type: 

 America borealis, Muhlenberg s.n. 

 (presumably B-WI, n.v.).— Hedyotis exigula 

 W. H. Lewis 



Houstonia longipes S. Watson, Proc. Amer. 

 Acad. Arts 18: 97. 1883. Syntypes: MEXICO: 

 Without precise locality, States of Coahuila 

 and Nuevo Leon, Feb-Oct 1880, E. Palmer 

 394 (GH, n.v.; US 47845 (photos MEXU, 

 PTBG)); Nuevo Leon, Monterrey, Feb 1-26 

 1880, E. Palmer 395 (GH, n.v., US 70739 

 (photos MEXU, PTBG), US 1390542 (photos 

 MEXU, PTBG)).— Hedyotis palmeri (A. 

 Gray) W. H. Lewis 



Houstonia mucronata (Benth.) B. L. Rob., Proc. 

 Amer. Acad. Arts 45: 401. 1910. Hedyotis 

 mucronata Benth., Bot. Voy. Sulphur 19. 

 1844. Type: MEXICO: Baja California, 

 Magdalena Bay, Barclay s.n. (Holotype BM, 

 n.v.). — Hedyotis mucronata Benth. 



Houstonia mullerae (Fosberg) Terrell, Brittonia 

 31: 169. 1979. Hedyotis mullerae Fosberg, 

 Lloydia 4: 288. 1941. Type: MEXICO: 

 Coahuila, Mun. de Cuatro Cienegas, Canon 

 del Agua, Sierra de la Madera, 9 Sep 1939, 

 C. H. Muller 3234 (Holotype US 2133305 ex 

 NA (photos MEXU, PTBG); Isotype GH, 

 n.v). — Hedyotis mullerae Fosberg 



Houstonia nigricans (Lam.) Fernald, Rhodora 

 42: 299. 1940. Gentiana nigricans Lam., 

 Encycl. 2: 645. 1786. Type: MEXICO: 

 Collector unknown, (presumably P-LAM, 

 n.v.).— Hedyotis nigricans (Lam.) Fosberg 



Houstonia palmeri A. Gray, Proc. Amer. Acad. 

 Arts 17: 202. 1882. Syntypes: MEXICO: 

 Coahuila, Sierra Madre, 40 mi. south of 

 Saltillo, 25-31 Jul 1880, E. Palmer 397 (GH, 

 n.v., US 47431 (photos MEXU, PTBG); E. 

 Palmer 398 (GH, n.v., F (2 sheets), US 70740 



