C. G. Pringle 6641 (Holotype GH, n.v.; 

 Isotypes A, n.v., MO, P (2 sheets; photos 

 MEXU, PTBG), US (photos MEXU, PTBG)). 

 — Machaonia acuminata Humb. & Bonpl. 



Machaonia galeottiana Baill., Bull. Mens. Soc. 

 Linn. Paris 1: 204. 1879. Type: CUBA: Le 

 Havane, H. Galeotti s.n. (Holotype P (photo 

 MO)). On the holotype label the word 

 "Veracruz" is crossed out and "Le Havane" 

 written in. As noted by Standley (1934: 222) 

 the collection resembles no Mexican species. 

 — Machaonia havanensis (Jacq.) Alain 



Machaonia hahniana Baill., Bull. Mens. Soc. 

 Linn. Paris 1: 204. 1879. Type: MEXICO: 

 Puebla, Acatlan, Juillet 1865, L. Hahn s.n. 

 (Holotype P (photos MEXU, MO, PTBG)). 



Machaonia havanensis (Jacq. ex J. F. Gmel.) 

 Alain, Phytologia 8: 370. 1962. Spermacoce 

 havanensis Jacq. in J. F. Gmel., Syst. Nat. 2: 

 234. 1791. Type: Based on Spermacoce 

 fruticosa Jacq., ined.? (type not located). 



Machaonia lindeniana Baill., Bull. Mens. Soc. 

 Linn. Paris 1: 204. 1879. Type: MEXICO: 

 Campeche, Campeche, fl. en Juin, (Yucatan), 

 Linden s.n. (Holotype P (photos MEXU, MO, 

 PTBG)). 



Machaonia martinezorum Lorence, Novon 4: 

 123, fig. 3. 1994. Type: MEXICO: Chiapas, 

 Mun. de Ocosingo, a 50 km al S de Boca 

 Lacantum, 130 m, 24 Jun 1986, E. Martinez S. 

 19016 (Holotype PTBG 7356; Isotypes 

 MEXU, MO). 



Machaonia martinicensis (DC.) Standi., Publ. 

 Field Mus. Nat. Hist., Bot. Ser. 22: 193. 1940. 

 Tertrea martinicensis DC, Prodr. 4: 481. 

 1830. Type: MARTINIQUE: Richard s.n. 

 (Holotype P, n.v.). 



Machaonia martinicensis (DC.) Standi, var. 

 dodgei (Standi.) Standi., Publ. Field Mus. Nat. 

 Hist., Bot. Ser. 22: 193. 1940. Machaonia 

 rotundata Griseb. var. dodgei Standi., Publ. 

 Field Mus. Nat. Hist., Bot. Ser. 18: 1326. 

 1938. Type: COSTA RICA: Beach between 

 Rio Sandio and Rio Tigre, 1 May 1930, C. W. 

 Dodge 7752 (Holotype F 890888 (photo 

 PTBG); Isotype MO (photo PTBG)). 

 — Machaonia martinicensis (DC.) Standi. 



Machaonia pringlei A. Gray, Proc. Amer. 

 Acad. Arts 21: 380. 1886. Type: MEXICO: 

 Coahuila, limestone ledges, mountains, 

 Jimulco, 27 Apr 1885, C. G. Pringle 146 

 (Holotype GH, n.v.; Isotypes A, n.v., F 

 (photos MEXU, PTBG), P (photos MEXU, 

 PTBG), US (photos MEXU, PTBG)). 



Machaonia rotundata Griseb. var. dodgei 

 Standi., Publ. Field Mus. Nat. Hist., Bot. Ser. 

 18: 1326. 1938. Type: COSTA RICA: Beach 

 between Rio Sandoval and River Tigre, 1 May 

 1930, C. W. Dodge 7752 (Holotype F 890888 

 (photo PTBG); Isotype MO (photo PTBG)). 

 — Machaonia martinicensis (DC.) Standi. 



Machaonia tysonii Dwyer, Ann. Missouri Bot. 

 Gard. 53: 106. 1966. Type: PANAMA. Los 

 Santos, thicket near man-made dam, main 

 highway, Monagre Beach, J. D. Dwyer 5067 

 (MO, not located in 6/1996). Note: John 

 Dwyer (pers. comm. 24 March 1993) believes 

 citation of this collection was a typographical 

 error and the following collection actually 

 represents the holotype: PANAMA: Along 

 Quebrada Vital off Rio Chucanagre, km 16 de 

 Yavita, 4 Jul 1962, J. A. Duke 5067 (Holotype 

 MO 3725554 (photo PTBG)). In the Flora of 

 Panama Rubiaceae treatment, Dwyer (1980) 

 erroneously cited Dwyer 5061 as the holotype; 

 the latter specimen was not located at MO in 

 3/1993 or 6/1996— Machanonia acuminata 

 Humb. & Bonpl. 



Machaonia velutina M. Martens & Galeotti, 

 Bull. Acad. Roy. Sci. Bruxelles 1 1: 134. 1844. 

 Type: MEXICO: Veracruz, ravines de Puente 

 Nacional, pres Veracruz, H. Galeotti 7104 

 (Holotype BR, n.v.; Isotype P (photos MEXU, 

 MO, PTBG)).— Machaonia acuminata 

 Humb. & Bonpl. 



Machaonia veracruzeana Baill., Bull. Mens. 

 Soc. Linn. Paris 1: 204. 1879. Norn, superfl., 

 based on Machaonia velutina M. Martens & 

 Galeotti, 1844 —Machaonia acuminata 

 Humb. & Bonpl. 



Macrocnemum P. Browne, Hist. Jam. 165. 



1756. Type: Macrocnemum jamaicense L. 

 Macrocnemum candidissimum Vahl, Symb. 2: 



38, pi. 30. 1791. Type: TRINIDAD: Vicinia 



