.PI.: 553, 558. 1771 



Bias, El Llano-Carti Rd., km 16.7, 350 m, trail 

 west to waterfall 5 km from road, 16 Jun 

 1985, G. de Nevers & S. Charnley 5907 

 (Holotype MO 3623655 (photo PTBG)). 



Manettia barbata Oerst., Vidensk. Meddel. 

 Dansk Naturhist. Foren. Kjobenhavn 1852: 

 47. 1853. Type: COSTA RICA: In Monte 

 Aguacate, 2000 ft., Feb 1846, Oersted 11521 

 (Holotype C, n.v. (photos MO, US); Isotypes 

 K, n.v., US (2 sheets; photos MEXU, PTBG)). 



\Iunenni hiicataurcnsis Dwyer, Ann. Missouri 

 Bot. Gard. 67: 278. 1980. Type: PANAMA: 

 Bocas del Toro, along RR (railroad) near 

 station at Milla 10, 27 Jul 1971, T. B. Croat & 

 D. M. Porter 16334 (Holotype MO 2166138 

 (photo PTBG)).— Hedyotis lancifolia K. 

 Schum. 



Manettia costaricensis Wernham, J. Bot. 4 1 

 (suppl.): 38. 1919. Syntypes: COSTA RICA: 

 Tucurrique, deboisements de Las Vueltas, 

 650-700 m, Dec 1898, A. Tonduz 12873 (B, 

 presumably destroyed (photos F, MO; 

 fragment F), BM, n.v., US 1390734 (photos 

 MEXU, PTBG), US 354369). COSTA RICA: 

 Without precise locality, 3500 ft., Endres 2 

 (W, n.v. (fragment F)).— Manettia reclinata 

 L. 



Manettia estrellae Standi., J. Wash. Acad. Sci. 

 15: 6. 1925. Type: COSTA RICA: Cartago, 

 La Estrella, 26-27 Mar 1924, P. C Standley 

 39224 (Holotype US 1 153146 (photos 

 MEXU, PTBG)) —Manettia flexilis 

 Brandegee 



Manettia flexilis Brandegee, Univ. Calif. Publ. 

 Bot. 6: 196. 1915. Type: MEXICO: Chiapas, 

 Cerro del Boqueron, Jun 1914, C. A. Purpus 

 7118 (Holotype UC 174907, n.v.; Isotypes F, 

 GH, n.v., MO (photos MEXU, PTBG), US 

 (photos MEXU, PTBG)). 



Manettia hirtella M. Martens & Galeotti, Bull. 

 Acad. Roy. Sci. Bruxelles 11: 238. 1844. 

 Type: MEXICO: Oaxaca, cordillere orientale, 

 au bords des rivieres, dans les forets de 

 Tonaguia, 4000-5000 pieds, Jun 1840, H. 



Galeotti 2635 (Holotype BR, r 



Manettia hydrophila Dwyer, Ann. Missouri Bot. 

 Gard. 67: 280. 1980. Type: PANAMA: 

 Veraguas, mouth of Rio Concepcion, in 

 swamp, 4 Dec 1967, W. H. Lewis, T. B. Croat 

 & J. L. Hawker 2822 (Holotype MO 

 2268375).— Sipanea biflora (L. f.) Cham. & 

 Schltdl. 



Manettia liebmannii Standi., J. Wash. Acad. Sci. 

 17: 337. 1927. Type: MEXICO: Oaxaca, 

 Pelado [probably Cerro Pelon], Liebmann 

 11485 [=Liebmann Rubiaceae 147] (Holotype 

 C, n.v. (fragment US; photos MEXU, 

 PTBG)).— Hedyotis galeottii (M. Martens) 

 Terrell & Lorence 



Manettia longicalycina Dwyer & Lorence, 

 Novon 3: 61. 1993. Type: PANAMA: Code, 

 Alto Calvario, 800-900 m, 20 Apr 1977, /. P. 

 Folsom & Jaslon 2665 (Holotype MO 

 2608777 (photo PTBG)). 



Manettia longipedicellata C. M. Taylor, Novon 

 5: 202,.% 1A. 1995. Type: COSTA RICA: 

 Limon, Canton de Matina, Cordillera 

 Talamanca, 200 m, aguas abajo de la 

 confluencia de la quebrada Canabral con el no 

 Barbilla, 10°00'10"N, 83°25'30"W, 5 Nov 

 1988, G. Herrera 2290 (Holotype CR, n.v.; 

 Isotype MO 3649361). 



Manettia microphylla Lorence & Dwyer, 

 Novon 3: 60. 1993. Type: PANAMA: Darien, 

 Pirre Massif, Alturas de Nique, above Cana 

 Mine, 850-1 150 m, 2 Mar 1988, G. 

 McPherson 12189 (Holotype MO 3648261 

 (photo PTBG); Isotypes PTBG, SCZ). 



Manettia orbifera Wernham, J. Bot. 57 (suppl.): 

 41. 1919. Type: GUATEMALA: Retalhuleu, 

 Bernoulli & Cairo 1649 (Holotype K, n.v.). 

 —Manettia reclinata L. 



Manettia panamensis Duchass. & Walp., 

 Linnaea 23: 753. 1850. Type: PANAMA: 

 Duchassaing s.n. (presumably GOET, n.v.). 

 — Manettia reclinata L. 



Manettia reclinata L., Mant. PI. 2: 558. 1771. 

 Type: Not designated. Note: this species is 

 actually based on Colombian material seen by 

 Mutis, not Mexican material. Apparently 

 Linnaeus believed that Santa Fe de Bogota 



