V7 
of Marattia and Angiopteris. In Danaea and Kaulfussia'), it 
is simpler, but in these also the relation of the bundles is not 
easy to follow. The anatomy in the very young sporophyte 
has been studied only very incompletely. Jeffrey has described the 
arrangement of the vascular system in young plants of Danaea, 
but did not have the earlier stages. Brebner has given an ac- 
count of the young sporophyte in D. simplicifolia, and Farmer 
described the structure in the very young sporangium in Angi- 
opteris, but not in the older stages, and the writer in a paper 
on Marattia douglasii gave a brief account of the arrangement 
of the bundles in the young sporophyte. 
A median section of the young sporophyte in both Angiopteris 
and Kaulfussia (Figs. 4 and 5) shows it to be a much elongated 
body penetrating the prothallium above and below, and made 
up principally of the cotyledon which emerges above and the 
root which emerges below, the limits between the two organs 
not being clearly defined. The foot which is so conspicuous in 
the early stages of the embryo can no longer be clearly re- 
cognized, as it has been carried down by the growth of the 
root and practically forms part of the root cap which is finally 
destroyed, and the foot is merged so completely with the tis- 
sues of the root apex as to be quite unrecognizable. How much 
of the outer tissue of the central region of the embryo between 
the bases of the root and cotyledon may be properly assigned 
to the foot it is impossible to say, but these cells which are 
in contact with the prothallial tissue undoubtedly function as 
absorbing cells and physiologically at least may be said to 
represent the foot. 
The stem apex projects but little or not at all and occupies 
& small area at the base of the cotyledon. Its outer surface 
may be nearly horizontal or it may be strongly inclined, and 
there may usually be recognized a single large cell which is 
ee 
1) Jeffrey, E. C.: The Structure and Development of the Stem in Pteridophytes 
and Gymnosperms. Phil. Trans. Royal Soc. Series B., 195, pages 119—146. 1902. 
Kiihn: Untersuchungen iiber die Anatomie der Marattiaceen und anderer Gefiss- 
Kryptogamen. Flora, p. 72, 457. 1889. 
