18 
the initial cell. This is somewhat elongated with a truncate 
base, and in transverse section generally is approximately qua- 
drilateral, although it is somewhat variable in form and may 
approach the triangular outline found in the leptosporangiate 
ferns. Segments are cut off laterally and also from the base, 
and the lateral segments also contribute to the growth in length 
of the stem, which, however, remains short so that the leaves 
of the young plant are much crowded. At this stage no central 
cylinder can be made out in the stem, but all of the procambial 
tissue belongs either to the cotyledon or to the root. 
The base of the petiole of the cotyledon in Angiopteris is 
of nearly the same size as the upper portion, but in Kaulfussia 
there is a marked depression in front, so that a cavity is formed 
at the base of the leaf somewhat as in Ophioglossum or Isoetes, 
and the stem apex is almost concealed within this cavity. 
About the time the cotyledon and root emerge, the second 
leaf makes its appearance, and as soon as it is recognizable, 
there may be seen a strand of procambium which joins that of 
the first leaf above its junction with the root. 
The cotyledon in Kaulfussiajis oval in outline and the venation 
much resembles that of Ophioglossum. In Angiopteris there is 
some difference. Farmer states that in the specimens examined 
by him there was a midrib in the cotyledon with lateral veins. 
I examined a considerable number of specimens and found that. 
while this type of cotyledon was sometimes met with, more 
commonly it approached the fan-shaped form of Marattia with 
dichotomous venation. There were all transitions between the 
strictly dichotomous venation. There were all transitions betweet 
the strictly dichotomous venation found in Marattia and the 
pinnate venation described by Farmer. 
The structure of the petiole is much alike in all of the forms. 
The ground tissue consists of the solid parenchyma and the 
vascular bundle which is nearly centrally placed has no evident 
endodermis and is concentric in structure, although the phloe™ 
is rather better developed on the outer side. In Kaulfussia 0° 
tannin cells are usually found in the cotyledon, but in Angi 
