79 
opteris these are very conspicuous and are found in immediate 
contact with the vascular bundle (Fig. 12). In the later leaves 
these tannin cells become very conspicuous and there are also 
formed large mucilage ducts. In both genera the root is diarch 
and there is a conspicuous endodermis with the characteristic 
thickening of the radial cells. The first two leaves in Angiopteris 
are destitute of the stipules which are so conspicuous in the 
later leaves, these first appearing in the third leaf. In Kaul- 
fussia the second leaf may develop stipules. 
The further study of the young sporophyte was confined to 
Kaulfussia, and it remains to be seen how far the later de- 
velopment of the sporophyte in the other genera agrees with this. 
The anatomy of the adult sporophyte in Kaulfussia has been 
carefully studied by Kuhn’), but he made no investigation 
upon the younger stages. The vascular system in Kaulfussia 
is much simpler than that of the other genera, and consists 
of the tubular reticulate cylinder with a single medullary strand. 
The stem in Kaulfussia is markedly dorsiventral and this be- 
comes evident in the very young sporophyte. Indeed, the po- 
sition of the second leaf, which is not opposite the cotyledon 
but turned toward the same side of the stem, indicates this, 
and by the time that three or four leaves are fully developed, 
the dorsiventral character of the sporophyte is very apparent. 
In the young sporophyte, at the time that it emerges from 
the prothallium, there is a single nearly axial vascular bundle 
which, as we have seen, belongs partly to the cotyledon, partly 
‘to the root, the line between being quite impossible to deter- 
mine, as the sections of the leaf bundle and that of the root 
are so much alike. 
Very soon after the first rudiment of the second leaf is vi- 
sible, as we have already indicated, a strand of procambium 
cells may be seen connecting it with the central vascular bundle 
at a point apparently near the junction of the primary root 
and leaf. No procambium could be demonstrated which seemed 
1) Loc. cit, 
