8] 
SUMMARY. 
1. It is probable that in all Marattiaceae stem, leaf and 
root all arise from the epibasal part of the embryo; in Danaea 
the primary hypobasal cell forms a suspensor, and all of the 
organs, including the foot are of epibasal origin. 
2. In Kaulfussia and Angiopteris the stem and leaf do not 
take up all of the epibasal tissue; a portion also contributes 
to the foot. 
3. The stem apex usually shows a single, large initial cell, 
which possibly may be derived directly from one of the primary 
quadrants of the embryo, the cotyledon cannot certainly be 
traced back to one of the primary quadrants and does not 
seem to be always formed from the same portion of the em- 
bryo; the cotyledon does not show a definite apical cell. 
4, The root arises secondarily, and is endogenous in origin; 
it is practically always of epibasal origin. It grows from a single 
initial cell. 
5. The foot is very large in the young embryo, but later 
is almost obliterated by the growth of the root. 
6. No vascular cylinder is formed in the stem region of the 
young sporophyte; the vascular bundle of the leaf is continued 
without interruption with the root, and the young sporophyte 
appears bipolar, the leaf growing upward, the root downward. 
7. The bundle of the leaf petiole is concentric; that of the 
root diarch. 
8. The vascular system of the stem of the young sporophyte 
is built up of the leaf traces alone. No strictly cauline bundles 
occur in the early stages of the spades 
Stanford University 
July 1909. 
Ann. Jard. bot. Buitenz. 2¢ Sér. Supplem. III. . 
