154 
UNGULATES. 
the third and fourth toes become symmetrical to a vertical line drawn between 
them. When this takes place the first toe disappears, and the second and fifth 
become diminished in size; an instance of this stage of development being presented 
BONES OF FORE- 
FOOT OF HORSE, 
by the pig, where the two large and medially - symmetrical toes 
represent the third and fourth of the typical series, while the 
two small lateral ones are the second and fifth. In the pigs! 
all the metacarpal bones remain distinct and relatively short; but 
in the water-chevrotain! of Africa the third and fourth meta- 
carpals become much elongated and closely appled to one another, 
while the second and fifth are reduced to mere splints, and their 
toes so diminished as to become practically functionless. Finally, 
in the deer, oxen, and their allies, the third and fourth metacarpals 
in the fore-limb, and the corresponding metatarsal bones in the 
hind-limb, have become completely fused into a single rod-like bone, 
corresponding in function with the cannon-bone of the horse, and 
generally known by the same name. The dual origin of this 
cannon-bone is, however, proclaimed by the formation of its lower extremity, 
which carries two pulley-like surfaces, with which the bones of the two functional 
toes (the third and fourth) articulate. 
BONES OF THE FORE AND HIND- 
EXTINCT DEER. 
FEET OF AN 

Since all the Ungulates displaying this second modi- 
fication of foot-structure agree in having the third and 
fourth toes arranged symmetrically to a line drawn 
between them, they are collectively termed the Even- 
Toed Ungulates. 
It is accordingly evident that although a few living 
Ungulates, like the elephant and the hyrax, retain a 
generalised type of foot, the greater number of the 
living representatives of the order are characterised by 
their more or less markedly specialised feet. 
As regards their teeth, the Ungulates 
are characterised by those of the cheek- 
series having broad crowns, surmounted either by columns 
or transverse ridges, and adapted for grinding and 
masticating vegetable substances. In the more specialised 
forms, like cattle and horses, these cheek-teeth have their 
columns or ridges of great height and closely approxi- 
mated to one another, in consequence of which the bases 
of the hollows, or valleys by which these columns or 
ridges are separated from one another, cannot be seen 
when the tooth is unworn; while the pattern produced 
on the crown by the wearing down of these columns or 
ridges is complex. On the other hand, in the more 
primitive types, such as pigs and tapirs, the crowns of the 
cheek-teeth have low columns, or ridges, so that the bases 
Teeth. 


—After Osborn, of the intervening valleys can be distinctly seen at all 
1 Figures of the bones of the feet of these animals are given under their respective headings. 
