) 



I 



1907] JEFFREY— ARAUCARIOPITYS 437 



canals of other Araucarian lignites from various levels of the 

 Cretaceous.^ 



Fig, 6 shows a tangential longitudinal section of the wood of our 

 species, through the region of the traumatic resin canals. It is to 

 be noted that these are more numerous than is the case with normal 

 resin canals, and are somewhat constricted at intervals, both features 

 to be found in the wound-canals of the Abietineae. In the region of 

 disturbance shoAvn in fig. 6^ the rays vary greatly in height, being 

 composed of from one to many cells. Fig. 7 shows the left-hand 

 portion of fig. 6, more highly magnified. In this figure the end 

 walls of the ray cells, which appear in face view, arc strongly pitted, 

 a feature found in no other living or fossil Araucarian, which has 

 been described, but very characteristic of the Abietineae. Both the 

 end and horizontal walls of the ray cells in Agathis and Araucaria, 

 for example, are smooth, and this statement equally holds for the 

 numerous species of ]\Iesozoic Araucarians which it has been possible 

 to study in the Androvette deposits. Our present species differs, 

 then, from all other Araucarians, living or extinct, in the presence of 

 abundant pitting on the terminal and horizontal walls of the cells 

 of its medullary rays. The cells constituting the rays are entirely 

 parenchymatous and there are no tracheidal cells present, such as 

 occur in the rays of certain species of living Abietineae. Fig. g 

 shows with sufficient clearness the strongly pitted horizontal wall 

 of the ray cells in our species. 



In fig. 8 is seen a longitudinal approximately radial section of 

 the wood in Araucariopitys. The magnification is scarcely sufficient 

 to show the araucarian radial pitting of the tracheids. This section 

 shows weiy well, however, the constriction of the traumatic resin 

 canals at intervals, which Professor Penhallow, in the case of the 

 traumatic and normal resin canals of certain Abietineae, has interpreted 

 as an indication of the derivation of the continuous systerri of resin 

 canals, present, for example, in Pinus, from the union with each other 

 of a number of originally isolated resin spaces or cysts.3 This view, 

 as will be shown on another occasion, can scarcely be accepted. 



M 



= Jeffrey, E. C-, The wound-reactions of Brach^-phyllum. Annals of Botany 



^o- 383-394. pis. 27, 28. 1906, 



3 Pexhallow, D. p., The anatomy of the Coniferales. Amer. Nat. 38:243 2"- 

 1904. 



