64 ADDISONIA 
1822, having a pouch-like lip with the opening in it turned down, 
two tail-like appendages, known as antennae, to the column, and 
the pollinia well-developed; this represents the staminate form. 
Another, almost exactly resembling this, has the pollinia imper- 
fectly developed, no appendages to the column, and represents the 
pistillate form; this was the basis of the genus Monachanthus, pub- 
lished by Lindley in 1832. Upon still a third form of flower Lindley 
in the same year established his genus Myanthus; in this the lip is 
flat, spreading, with a well-pronounced spur, and the column is 
provided with appendages; this represents the perfect form of 
flower. In 1836, however, a plant of Myanthus cristatus in 
the collection of the Duke of Devonshire put forth a cluster of 
flowers which shattered the opinions previously held by botanists. 
This flower-cluster had upon it flowers of all three forms, thus 
clearly demonstrating that the three genera were in reality only 
forms of one. ‘The presence of the appendages to the column in 
the perfect and staminate forms makes quite evident that these 
organs play an important part in the fertilization of the plant. An 
irritation of these appendages results in an immediate and violent 
expulsion of the pollinia, which fasten themselves, by means of the 
sticky gland at the end of the stalk, to the first object they 
encounter. Cycnoches, belonging to the same tribe and closely related 
to Catasetum, exhibits a similar variability in flower-form. 
GEORGE V. Nasu. 
EXPLANATION OF PLATE. Fig. 1.—Plant. Fig. 2.—Vertical section of 
Fig. 4.— 
flower, somewhat enlarged. Fig. 3.—Column, front view, X 2. 
Pollinia, front view, X 4. Fig. 5.—Pollinia, dorsal view, X 4 
