VOLUME XLIII NUMBER 1 
BOTANICAL. GAZETTE 
JANUARY 17907 
FERTILIZATION AND EMBRYOGENY IN CEPHALO- 
TAXUS FORTUNEI 
W:; C. CoEER 
(WITH FIVE FIGURES AND PLATE I) 
The material for this study was collected at Darlington, S. C., 
where the plant grew normally and produced good seeds. Collec- 
tions of young ovules were also made in Bonn, Germany, but they 
are here represented only in jig. 1. 
Our knowledge of the spermatogenesis of Cephalotaxus is confined 
to the work of SrrasBuRGER (18) and of ARNOLDI (1). Twenty- 
seven years ago STRASBURGER described the development of the 
embryo quite accurately for the stages observed, but he did not secure 
young proembryos, ARNotpr has described the gametophytes and 
proembryo, but he has overlooked certain interesting peculiarities 
and is in places not sufficiently clear. 
Material is not yet at hand to determine the development of the 
young ovule in detail, and its early history will not be considered 
here. However, one figure is given (fig. 1) from material collected 
January 5, 1902, at Bonn, to show the interesting midwinter condi- 
tion of the ovule. This is about nine and one-half months after 
pollination and the pollen tube has developed into a large sac which 
occupies a great part of the tip of the nucellus. The body cell and 
two vegetative nuclei are noticed near the center. In the massive 
lower part of the ovule is the megaspore, not yet divided. This 
winter condition will at once suggest the great difference between 
the genera of the Taxeae in regard to the time elapsing between the 
critical points of pollination, fertilization, and maturation of the seeds, 
In both Torreya (Courter and Lan, 6) and Cephalotaxus the 
: I 
