1907] COKER—CEPHALOTAXUS FORTUNEI 9 
in walls and may divide after their formation, In one figure he shows 
each rosette cell divided into a group. According to him the embryo 
proper is formed from the group of cells lying between the tip cell 
(or cells) and the suspensors, and the tip cell is destroyed. 
In comparing the proembryo of Cephalotaxus with that of its 
relatives, we find that it shows some resemblance to Taxus, but 
very little to Torreya. In Taxus, according to JAEGER (8), cell 
walls are not formed before the sixteen cell-stage at the earliest, but, 
as one would expect from the shape of the archegonium, there is no 
long and narrow cell at the apex of the embryo. In Torreya, accord- 
ing to Miss RoBERTSON (17) and CouLter and Lanp (6), cell walls 
are formed in the four cell-stage. In Torreya californica the organized 
proembryo consists of an exposed rosette, a tier of four or six sus- 
pensors, and a “cluster” of tip cells (Miss RoBertson, 17). In 
T. taxifolia, however, the proembryo entirely fills the archegonium 
with twelve or eighteen cells, all closed, and in this stage passes the 
winter (COULTER and Lanp, 6). Cephalotaxus, Taxus, and Podo- 
carpus are the only conifers so far examined in which cell walls are 
not formed in the proembryo before the sixteen cell-stage. 
Cuaper Hirt, N. C. 
LITERATURE CITED 
1. ARNotpDr, W., Beitriige zur Morphologie der Gymnospermen, III. Flora 
87: 46-63. ae I~}. 1900. 
oat Beitriige zur MMospiiclanie der Gymnospermen, V. Bull. Soc. Nat. 
Moscow no. 4. pp. 28. pls. 2. 1900. 
3. BELayerr, W. C., Zur Lehre von dem Pollenschlauche der Gymnospermen. 
Ber. Deutsch. Bot. Gesells. 9: 280-286. pl. 18. 1891. 
- Coxer, W. C., Notes on the Papago and embryo of Podocarpus. 
Bor. Gazerre 33: 89-107. 
S- ———On the gametophytes Sa ved ee Taxodium. Bor. GAZETTE 
36:1-27, 114-140. pls. 1-11. 1903. 
- CouLrer, Joun M. and Lanp, W. J. G., Gametophytes and embryo of 
Torreya taxifolia. Bor. GAZETTE 39:161~-178. pls. A and 1-3. 1905. 
- Fercuson, Marcaret C., The development of the pollen tube and the 
division of the generative nucleus in a species of Pinus. 
of Botany 15: 193-223. pls. 12-14. 1 
8. Jarcer, L. , Beitrige zur Kenntniss der Pakopest iin und zur Embry- 
ologie von Taxus baccata L. Flora 86: 241-288. pls. 15-19. 1899. 
9. Lanp, W.J.G.,A morphological study of Thuja. Bot. GAZETTE 34:249- 
259. pls. 6-8. go2. 
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