126 BOTANICAL GAZETTE [FEBRUARY 
remaining layers extend through its entire length, being the same 
both where it is separate from the nucellus and where it is continuous 
with it. The outer of these three layers becomes the fleshy, juicy part of 
the ripened fruit (figs. 34, 35); the middle layer, by a great thickening 
of walls and compacting of cells, becomes the stony coat; the inner 
fleshy layer, which is delicate and watery in early stages, becomes 
crushed and dry, forming the papery layer which lines the stony coat 
and adheres to it in the mature fruit. The nucellar tissue is quite 
distinct from the watery tissue of the integument, and in the upper 
part the two are not even in contact (fig. 4o), so the former cannot 
form the papery coat as it is said to do by Sewarp and GowaN (9). 
Below the line of junction of the nucellus and integument the former 
may contribute some small part to the papery coat, but it is almost 
entirely absorbed by the prothallium. 
At the base of the ovule two bundles enter the watery tissue, coming 
up through a little gap in the tissue of a stony coat (fig. go). Their 
course lies along the inner surface of the watery tissue, one curving up 
on each side and ending just below the point at which the integument 
becomes free from the nucellus. The abundant liquid in this watery 
tissue doubtless comes up through these bundles. Upon removal 
of the integument, which splits readily into two lips, the nucellus is 
seen to be flattened on two opposite sides, the intervening sides being 
angled (figs. 34, 35). It is in these angled sides that the bundles end 
(jig. 35). In some cases there is a three-lipped integument and a 
three-angled nucellus, there being also three bundles, one ending in 
each angled side. 
SUMMARY 
1. The ovule has a conspicuous nucellar beak and pollen chamber. 
2. The sporogenous tissue is deep within the nucellus. 
3. Differentiation of the mother cell has taken place by the first. 
of May. 
4. One mother cell is usual, but more may occur. 
5. A peculiar kinoplasmic mass is present in the mother cell. 
6. The gametophyte number of chromosomes is eight. 
7. The first spindle is obliquely placed i in ~ mother cell. 
8. The tetrad may be complete or 
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