166 BOTANICAL GAZETTE [MARCH 
tains raphides but no starch. This same structure is to be observed 
in the upper internodes except that the outline becomes more sharply 
four-winged as the cortical parenchyma in the wings becomes collen- 
chymatic, and the hairs are more abundant. 
The peduncle of the flower, however, is not quadrangular, but 
cylindric, and the hairs are reduced to mere papillae, which are 
exceedingly numerous. There is no stereome and no collenchyma; 
the cortical parenchyma is thin-walled, very compact, and the endo- 
dermis consists of much larger cells than observed in the internodes. 
The structure of the central cylinder is the same. 
The leaves.—The over-wintering leaves are petiolate; the petiole 
is triangular in outline and covered by a wrinkled cuticle. The epi- 
dermis is moderately thickened on the ventral face, but thin-walled 
on the dorsal. A thin-walled chlorenchyma with very little chloro- 
phyll and raphides surrounds three veins, the median of which is the 
largest; it is crescent-shaped and contains a collateral mestome 
strand surrounded by a thin-walled, colorless parenchyma sheath. 
The lateral veins are much smaller and round in transverse section. 
The blades of these basal leaves are sparingly hairy on the upper 
face from short, unicellular, obtuse hairs with very distinct longi- 
tudinal ridges of cuticle; but where the cuticle covers the ordinary 
cells of the epidermis (not the hairs) on the upper face we meet with 
a corresponding striation as observed in H. coerulea (fig. gA). The 
lateral cell walls of the epidermis are undulate on both faces, especially 
on the lower, and it seems characteristic of this species (when com- 
pared with H. coerulea) that the lateral walls show local thickenings 
(figs.g and 9A). This structure I observed, also, and much farther 
developed in Mitchella, where it will be described more particularly. 
Stomata occur only on the lower face and show the same structure 
as those of H. coerulea. There is a typical palisade tissue of two 
layers, and a very open pneumatic tissue with the cells oblong and 
parallel with the leaf surface. The mestome strands are surrounded 
by parenchyma sheaths, but have no mechanical support, neither of 
collenchyma nor of stereome. 
The stem leaves are sessile and their structure differs somewhat 
from that of the basal leaves. For instance, the cuticle does not 
show the stellate striations, and bicellular hairs (fig. 8) abound on 
