418 BOTANICAL GAZETTE [JUNE 
together under proper nutrient conditions. The form is therefore hetero- 
thallic. 
Whether or not it may ever be found that a homothallic race may occur 
in a species normally heterothallic (perhaps not an impossible condition in 
view of the writer’s having obtained a homothallic mycelium of the hetero- 
thallic Phycomyces), the evidence at hand leads one to the conclusion that 
the large majority of the Mucorineae are heterothallic.—A. F. BLAKESLEE, 
Cryptogamic Laboratory, Harvard University. 
TWO EMBRYO-SAC MOTHER CELLS IN LILIUM 
LONGIFLORUM ; 
(WITH ONE FIGURE) 
The presence of more than one archesporial cell in the megasporangium 
of the Spermatophyta has frequently been reported for gymnosperms and 
dicotyledons, but the records of their occurrence in monocotyledons are 
very few. In 1882 GUIGNARD* 
figured two small hypodermal 
cells in Ornithogalum pyrena- 
icum, which he interprets as 
archesporial. As these cells 
are but little larger than the 
other cells of the nucellus, and 
the character of the proto- 
plasmic content of the adjacent 
cells is not shown, the evidence 
here is not altogether conclu- 
sive. CovuLTER and CHAM- 
BERLAIN? state that they have 
observed two preparations of 
Lilium philadelphicum show- 
Fic. 1.—Vertical section through a young jing respectively three and five 
ovale of Lilium longiflorum, showing two archesporial cells, but they 
aa aera : neither described nor figured 
them. BERNARD? observed and figured two four-celled embryo sacs 
* Guicnarp, L., Recherches sur le sac embryonnaire des Phanérogames Angio- 
spermes. Ann. Sci. Nat. Bot. VI. 13:1 36-199. pls. 3-7. 1882. 
? CoutTER and CHAMBERLAIN, Morphology of Angiosperms. New York. 1903- 
3 BERNARD, C. H., Recherches sur les sphtres attractives chez Lilium candidum, 
etc. Jour. Botanique 14:178. pls. 4-5. 1900. 
