1888. ] BOTANICAL GAZETTE. 207 
0, Hemerocallis, Pontederta, etc., among Moffocotyle- 
ns. pene 
Some of these flowers show no indications of the condi- 
tions which are supposed to have produced them. Thus, 
Trifolium pratense is no longer shallow, but is deep and tubu- 
lar, fe insect no longer lighting on the flower, but on the 
head, and the pollen is dusted on the under side of the bee’s 
head or proboscis. My theory only requires what would be 
- lost wings and keel, only retaining the vexillum, seca 
makes the spikes more conspicuous, and its insect relation 
are much the same as in the apetalous Salix. poet: 
But there are some flowers of polypétalous Soe Viola 
are Nototribe, such as most orchids, Impatiens and V10 
_ (sometimes). % ibe, and 
-_ The orchids must have been developed as ste Hage rts the 
become nototribe by the twist in the ovary, which ive or- 
flower, The labellum must have been developed aber the 
82n of attraction, like the vexillum of Papilignacee 4 land- 
“Per colored sepals of Pontederia, and has become a It is 
& °0 account of the flower turning upside done: uired 
Temarkable that these flowers, whose structure ‘ea 
